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Home -> Kingdom Animalia -> Phylum Chordata -> Subphylum Vertebrata -> Class Mammalia -> Order Artiodactyla -> Family Cervidae -> Subfamily Cervinae -> Species Rusa unicolor

Rusa unicolor
sambar



2009/11/08 05:02:54.695 US/Eastern

By Christine Brown

Kingdom: Animalia
Phylum: Chordata
Subphylum: Vertebrata
Class: Mammalia
Order: Artiodactyla
Family: Cervidae
Subfamily: Cervinae
Genus: Rusa
Species: Rusa unicolor

Geographic Range

Rusa unicolor are native to India, Pakistan, Ceylon, Burma, Sri Lanka, Philippines, southern China, Taiwan, Malaysia, Borneo, Sumatra, and Java (Medway, 1969). Sambars have also been successfully introduced in Australia, New Zealand, California, Florida and Texas (Nowak, 1999). (Medway, 1969; Nowak, 1999)

Biogeographic Regions:
nearctic (introduced ); palearctic (native ); oriental (native ); australian (introduced ).

Habitat

Elevation
3700 m (high)
(12136 ft)


Sambars inhabit both gently sloping and steep forested hillsides. They reside preferably near cultivated areas, such as gardens and plantations, in order to acquire food, but are also found in thick forests, swamp forests, and open scrub (Payne et al., 1985). Some of these deer may move between higher altitudes in the summer to lower, more sheltered areas during the winter months (Nowak, 1999). (Nowak, 1999; Payne, Francis, and Phillipps, 1985)

These animals are found in the following types of habitat:
temperate ; tropical ; terrestrial .

Terrestrial Biomes:
forest ; scrub forest .

Wetlands: swamp .

Other:
agricultural .

Physical Description

Mass
109 to 260 kg; avg. 184.50 kg
(239.8 to 572 lbs; avg. 405.9 lbs)


Length
1620 to 2460 mm; avg. 2040 mm
(63.78 to 96.85 in; avg. 80.31 in)


Sambars have a coarse coat of short, dark hair with lighter brown to creamy white hair on their undersides (Tate, 1947). The backsides and undersides of their bushy tails are white, and when raised, the tails are used as signals (Nowak, 1999). Males are generally larger than females and posess a dense mane on their necks. Male Sambars have antlers with three or four tines, and these antlers are periodically shed and replaced. Antlers can reach lengths of up to 100 cm. The maximum size for males is 185-260 kg, and about 162 kg for females (Medway, 1969). (Medway, 1969; Nowak, 1999; Tate, 1947)

Some key physical features:
endothermic ; homoiothermic; bilateral symmetry .

Reproduction

Breeding interval
Sambars breed once yearly.

Breeding season
Sambars have no specific breeding season, but breeding most commonly occurs from September through January.

Number of offspring
1 to 2; avg. 1

Gestation period
8 to 9 months; avg. 8.50 months

Time to weaning
12 to 24 months; avg. 18 months

Age at sexual or reproductive maturity (female)
2 years (average)

Age at sexual or reproductive maturity (male)
2 years (average)

Male Sambars are solitary and very agressive during the breeding season, while females may be found in groups of up to 8 individuals (Nowak, 1999). Their mating system is polygynous, with one male mating with as many females as he can. (Nowak, 1999)

Mating systems:
polygynous .

Sambars have no specific breeding season, but breeding most commonly occurs from September through January. Usually only one fawn is born at a time, and the gestation period is about 9 months. (Medway, 1969). At birth, Rusa unicolor are very active and have brown hair with lighter spots, which are soon lost shortly (Medway,1969). Fawns weigh about 10 kg at birth. Males develop small antlers at one to two years; at three years antlers have two points, and adult males have antlers with three or four points (Payne et al., 1985). Females reach sexual maturity at approximately two years of age (Nowak, 1999). Males are aggressive towards one another during the mating season and actively defend territories, mating with multiple females that enter the area. Males mark their territory with scent glands, and as many as 8 females at a time may remain with one male within his range (Nowak, 1999). (Medway, 1969; Nowak, 1999; Payne, Francis, and Phillipps, 1985)

Key reproductive features:
iteroparous ; year-round breeding ; gonochoric/gonochoristic/dioecious (sexes separate); sexual ; viviparous .

Fawns are born weak but able to walk. They remain with their mother for 1-2 years. (Medway, 1969)

Parental investment:
precocial ; pre-fertilization (provisioning, protecting: female); pre-hatching/birth (provisioning: female, protecting: female); pre-weaning/fledging (provisioning: female, protecting: female); pre-independence (protecting: female); post-independence association with parents.

Lifespan/Longevity

Extreme lifespan (captivity)
26 years (high)

Average lifespan (wild)
20 years

In captivity, Rusa unicolor can live for over 26 years, but in the wild the average lifespan is about 20 years (Nowak, 1999). (Nowak, 1999)

Behavior

Sambars are mostly nocturnal, resting during the day in heavy forest cover (Medway, 1969). These deer are solitary but may be found in small groups during the mating season. Also, groups of up to 6 females with dependent young may travel together (Payne et al., 1985). Males are nomadic and establish territories primarily during breeding seasons. All sambars are proficient swimmers (Nowak, 1999). (Medway, 1969; Nowak, 1999; Payne, Francis, and Phillipps, 1985)

Key behaviors:
cursorial; terricolous; nocturnal ; crepuscular ; motile ; nomadic ; solitary ; territorial ; social .

Food Habits

Sambars generally feed at dusk or at night, and they browse on leaves, berries, grasses, bark from young trees, fallen fruit, herbs and buds. They browse mainly at clearings and forest edges (Payne et al., 1985). (Payne, Francis, and Phillipps, 1985)

Primary Diet:
herbivore (folivore ).

Plant Foods:
leaves; wood, bark, or stems; fruit.

Predation

Known predators

Sambars have developed a crepuscular and nocturnal activity pattern in response to hunting pressures from humans (Payne et al., 1985). (Payne, Francis, and Phillipps, 1985)

Ecosystem Roles

Sambar may disperse seeds as they forage.

Key ways these animals impact their ecosystem:
disperses seeds.

Economic Importance for Humans: Negative

Sambars may damage agricultural crops when they forage near human habitation.

Ways that these animals might be a problem for humans:
crop pest.

Economic Importance for Humans: Positive

Humans hunt Rusa unicolor for food and for trade. Sambars are also captured and placed into zoos worldwide. (Payne, Francis, and Phillipps, 1985)

Ways that people benefit from these animals:
food ; body parts are source of valuable material; ecotourism .

Conservation Status

IUCN Red List: [link]:
Lower Risk - Least Concern.

US Federal List: [link]:
No special status.

CITES: [link]:
No special status.

Although global population data is not known, the population in India exceeds 50,000 and in Australia Sambars number more than 5,000 individuals. (Nowak, 1999)

Other Comments

Several subspecies are recognized, including Rusa unicolor equinus, Rusa unicolor unicolor, Rusa unicolor brookei, and Rusa unicolor dejeani. Sambars were previously considered a member of the genus Cervus, as Cervus unicolor. (Payne, Francis, and Phillipps, 1985; Tate, 1947)

For More Information

Find Rusa unicolor information at

Contributors

Christine Brown (author), University of Michigan.
Ondrej Podlaha (editor), University of Michigan.

References

CITES Secretariat, 2001. "Convention on International Trade in Endangered Species of Wild Fauna and Flora" (On-line). Accessed November 18, 2001 at www.cites.org.

Medway, L. 1969. The Wild Mammals of Malaya. London: Oxford University Press.

Nowak, R. 1999. Walker's Mammals of the World, 6th ed.. Baltimore and London: The Johns Hopkins University Press.

Payne, J., C. Francis, K. Phillipps. 1985. A Field Guide to the Mammals of Borneo. Malaysia: The Sabah Society with WWF Malaysia.

Tate, G. 1947. Mammals of Eastern Asia. New York: The Macmillan Company.

The World Conservation Union, 2000. "2000 IUCN Red List of Threatened Species" (On-line). Accessed November 18, 2001 at www.redlist.org.

2009/11/08 05:02:56.224 US/Eastern

To cite this page: Brown, C. 2002. "Rusa unicolor" (On-line), Animal Diversity Web. Accessed November 09, 2009 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Rusa_unicolor.html.

Disclaimer: The Animal Diversity Web is an educational resource written largely by and for college students. ADW doesn't cover all species in the world, nor does it include all the latest scientific information about organisms we describe. Though we edit our accounts for accuracy, we cannot guarantee all information in those accounts. While ADW staff and contributors provide references to books and websites that we believe are reputable, we cannot necessarily endorse the contents of references beyond our control.

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