By Christine Brown
Geographic Range
Rusa unicolor are native to India, Pakistan, Ceylon, Burma, Sri Lanka, Philippines, southern China, Taiwan, Malaysia, Borneo, Sumatra, and Java (Medway, 1969). Sambars have also been successfully introduced in Australia, New Zealand, California, Florida and Texas (Nowak, 1999). (Medway, 1969; Nowak, 1999)
Biogeographic Regions:
nearctic
(introduced
); palearctic
(native
); oriental
(native
); australian
(introduced
).
Habitat
(12136 ft)
Sambars inhabit both gently sloping and steep forested hillsides. They reside preferably near cultivated areas, such as gardens and plantations, in order to acquire food, but are also found in thick forests, swamp forests, and open scrub (Payne et al., 1985). Some of these deer may move between higher altitudes in the summer to lower, more sheltered areas during the winter months (Nowak, 1999). (Nowak, 1999; Payne, Francis, and Phillipps, 1985)
These animals are found in the following types of habitat:
temperate
; tropical
; terrestrial
.
Terrestrial Biomes:
forest
; scrub forest
.
Wetlands: swamp
.
Other:
agricultural
.
Physical Description
(239.8 to 572 lbs; avg. 405.9 lbs)
(63.78 to 96.85 in; avg. 80.31 in)
Sambars have a coarse coat of short, dark hair with lighter brown to creamy white hair on their undersides (Tate, 1947). The backsides and undersides of their bushy tails are white, and when raised, the tails are used as signals (Nowak, 1999). Males are generally larger than females and posess a dense mane on their necks. Male Sambars have antlers with three or four tines, and these antlers are periodically shed and replaced. Antlers can reach lengths of up to 100 cm. The maximum size for males is 185-260 kg, and about 162 kg for females (Medway, 1969). (Medway, 1969; Nowak, 1999; Tate, 1947)
Some key physical features:
endothermic
; homoiothermic; bilateral symmetry
.
Sexual dimorphism:
male larger, ornamentation
.
Reproduction
Sambars breed once yearly.
Sambars have no specific breeding season, but breeding most commonly occurs from September through January.
Male Sambars are solitary and very agressive during the breeding season, while females may be found in groups of up to 8 individuals (Nowak, 1999). Their mating system is polygynous, with one male mating with as many females as he can. (Nowak, 1999)
Mating systems:
polygynous
.
Sambars have no specific breeding season, but breeding most commonly occurs from September through January. Usually only one fawn is born at a time, and the gestation period is about 9 months. (Medway, 1969). At birth, Rusa unicolor are very active and have brown hair with lighter spots, which are soon lost shortly (Medway,1969). Fawns weigh about 10 kg at birth. Males develop small antlers at one to two years; at three years antlers have two points, and adult males have antlers with three or four points (Payne et al., 1985). Females reach sexual maturity at approximately two years of age (Nowak, 1999). Males are aggressive towards one another during the mating season and actively defend territories, mating with multiple females that enter the area. Males mark their territory with scent glands, and as many as 8 females at a time may remain with one male within his range (Nowak, 1999). (Medway, 1969; Nowak, 1999; Payne, Francis, and Phillipps, 1985)
Key reproductive features:
iteroparous
; year-round breeding
; gonochoric/gonochoristic/dioecious (sexes separate); sexual
; viviparous
.
Fawns are born weak but able to walk. They remain with their mother for 1-2 years. (Medway, 1969)
Parental investment:
precocial
; pre-fertilization (provisioning, protecting: female); pre-hatching/birth (provisioning: female, protecting: female); pre-weaning/fledging (provisioning: female, protecting: female); pre-independence (protecting: female); post-independence association with parents.
Lifespan/Longevity
In captivity, Rusa unicolor can live for over 26 years, but in the wild the average lifespan is about 20 years (Nowak, 1999). (Nowak, 1999)
Behavior
Sambars are mostly nocturnal, resting during the day in heavy forest cover (Medway, 1969). These deer are solitary but may be found in small groups during the mating season. Also, groups of up to 6 females with dependent young may travel together (Payne et al., 1985). Males are nomadic and establish territories primarily during breeding seasons. All sambars are proficient swimmers (Nowak, 1999). (Medway, 1969; Nowak, 1999; Payne, Francis, and Phillipps, 1985)
Key behaviors:
cursorial; terricolous; nocturnal
; crepuscular
; motile
; nomadic
; solitary
; territorial
; social
.
Food Habits
Sambars generally feed at dusk or at night, and they browse on leaves, berries, grasses, bark from young trees, fallen fruit, herbs and buds. They browse mainly at clearings and forest edges (Payne et al., 1985). (Payne, Francis, and Phillipps, 1985)
Plant Foods:
leaves; wood, bark, or stems; fruit.
Predation
- humans (Homo sapiens)
- tigers (Panthera tigris)
Sambars have developed a crepuscular and nocturnal activity pattern in response to hunting pressures from humans (Payne et al., 1985). (Payne, Francis, and Phillipps, 1985)
Ecosystem Roles
Sambar may disperse seeds as they forage.
Key ways these animals impact their ecosystem:
disperses seeds.
Economic Importance for Humans: Negative
Sambars may damage agricultural crops when they forage near human habitation.
Ways that these animals might be a problem for humans:
crop pest.
Economic Importance for Humans: Positive
Humans hunt Rusa unicolor for food and for trade. Sambars are also captured and placed into zoos worldwide. (Payne, Francis, and Phillipps, 1985)
Ways that people benefit from these animals:
food
; body parts are source of valuable material; ecotourism
.
Conservation Status
IUCN Red List: [link]:
Lower Risk - Least Concern.
US Federal List: [link]:
No special status.
CITES: [link]:
No special status.
Although global population data is not known, the population in India exceeds 50,000 and in Australia Sambars number more than 5,000 individuals. (Nowak, 1999)
Other Comments
Several subspecies are recognized, including Rusa unicolor equinus, Rusa unicolor unicolor, Rusa unicolor brookei, and Rusa unicolor dejeani. Sambars were previously considered a member of the genus Cervus, as Cervus unicolor. (Payne, Francis, and Phillipps, 1985; Tate, 1947)
For More Information
Find Rusa unicolor information at
Contributors
Christine Brown (author), University of Michigan.
Ondrej Podlaha (editor), University of Michigan.

