Phocoena phocoenaharbor porpoise

Ge­o­graphic Range

Found in coastal re­gions of the North At­lantic, Arc­tic, and North Pa­cific Oceans; also the Mediter­ranean and Black Sea re­gions. They are found in bays, es­tu­ar­ies, river mouths, and some­times as­cend fur­ther up rivers. (Dollinger (ed­i­tor), 1988; Nowak, 1999; Dollinger (ed­i­tor), 1988; Nowak, 1999)

Habi­tat

Found in both salt and fresh water areas, Pho­coena pho­coena re­quire a shal­low coastal body of water. In the West­ern At­lantic, they also move far out to sea near the end of sum­mer and reap­pear in spring. Other re­gional pop­u­la­tions move south or far­ther away from shore to avoid ice buildups. (Nowak, 1999)

Phys­i­cal De­scrip­tion

Pho­coena pho­coena, also known as the "Com­mon" or "Har­bor" Por­poise, is a small cetacean that is 1.5 to 2 me­ters long and weighs from 45 - 65 kilo­grams. The fe­male of the species is usu­ally slightly larger than the male. The color of the an­i­mal varies from in­di­vid­ual to in­di­vid­ual, but the most com­mon col­oration pat­tern is a dark dor­sal sur­face that shifts to a lighter col­ored hue ven­trally. Al­though the dark color is usu­ally black or deep gray, al­bi­nos have been re­ported in which the dark seg­ments are com­pletely or par­tially white. The flip­pers, dor­sal fin, and tail are all dark in color, and there is a black stripe that runs from the edge of the mouth or eye to the flip­per on ei­ther side. There is no no­tice­able fore­head or beak on this species, and the snout is short, giv­ing the head a some­what cone-like shape. P. pho­coena has two pec­toral flip­pers, a sin­gle dor­sal fin, and a tail with two par­tially sep­a­rated flukes. All of these ap­pendages are short and not very sharp, with the dor­sal fin being tri­an­gu­lar shaped and usu­ally around 15 - 20 cm tall. There is a no­tice­able keel lo­cated near the all dark tail flukes, with the tail it­self span­ning any­where from 30-65 cm. In­side the slightly up­turned mouth there are rows of 16-28 spade-shaped teeth. There is no vari­ance in the shape or type of teeth in P. pho­coena. (Dollinger (ed­i­tor), 1988; Nowak, 1999; The Por­poise Page, 1998)

  • Sexual Dimorphism
  • female larger
  • Range mass
    45 to 60 kg
    99.12 to 132.16 lb
  • Range length
    1.5 to 2 m
    4.92 to 6.56 ft
  • Average basal metabolic rate
    107.675 W
    AnAge

Re­pro­duc­tion

It is not clearly known how ex­ten­sive the mat­ing sea­son is in P. pho­coena, but it seems that mat­ing mainly oc­curs from June to Sep­tem­ber with births oc­cur­ring from May to Au­gust. It is com­monly noted that ges­ta­tion lasts 11 months with nurs­ing fol­low­ing for an­other 7 or 8 months. A fe­male will give birth to one calf per year, with the birth size of the calf being 6-8kg and 70-100cm long. Sex­ual ma­tu­rity is reached by the fifth year, if not be­fore, and the life span of P. pho­coena is be­lieved to be any­where from 6 to 20 years. (John­ston(1999), Nowak(1999))

  • Breeding interval
    A female will give birth to one calf per year
  • Breeding season
    Mating mainly occurs from June to September
  • Average number of offspring
    1
  • Average number of offspring
    1
    AnAge
  • Average gestation period
    11 months
  • Average gestation period
    320 days
    AnAge
  • Range weaning age
    7 to 8 months
  • Range age at sexual or reproductive maturity (female)
    5 (high) years
  • Range age at sexual or reproductive maturity (male)
    5 (high) years

Lifes­pan/Longevity

Be­hav­ior

The species usu­ally swims near the sur­face, ris­ing to the sur­face to breath about every 25 sec­onds. It does not pre­sent a very play­ful at­ti­tude, ig­nor­ing boats and rarely jump­ing out of the water. Har­bor Por­poises do not move par­tic­u­larly fast, but when pur­sued can reach speeds of around 23km/hr. When div­ing for food this por­poise stays down for an av­er­age of 4 min­utes, and is be­lieved to be able to dive as deep as 200m. Al­though schools of up to 100 in­di­vid­u­als may some­times be seen, P. pho­coena is usu­ally seen in pairs or in groups of 5-10. When the larger groups do occur, it is usu­ally be­cause a num­ber of smaller groups have joined to­gether while fol­low­ing a rich food source. Moth­ers bring new­born calves to se­cluded coves to nurse. Some pop­u­la­tions are known to mi­grate, but when they re­turn to their reg­u­lar wa­ters they are ter­ri­to­r­ial, pa­trolling cer­tain areas. (John­ston(1999), Nowak(1999))

Com­mu­ni­ca­tion and Per­cep­tion

Food Habits

Diet con­sists mainly of smooth, non-spiny fish, and cephalopods. Her­ring, pol­lack, hake, sar­dines, and cod are com­monly eaten. Other sea crea­tures such as squid and shrimp are also con­sumed. P. pho­coena pro­duces click-like sounds sim­i­lar to those used by other cetaceans as a means of echolo­ca­tion in order to lo­cate food.(John­ston(1999), Nowak(1999))

  • Animal Foods
  • fish
  • mollusks
  • aquatic crustaceans

Eco­nomic Im­por­tance for Hu­mans: Pos­i­tive

Taken heav­ily in var­i­ous areas, the meat is used for human and an­i­mal con­sump­tion, and its oil is used in lamps and as a lu­bri­cant.

Con­ser­va­tion Sta­tus

Al­though fish­ing of P. pho­coena is now il­le­gal in most areas, the species is still in dan­ger. De­lib­er­ate and ac­ci­den­tal deaths still occur be­cause mod­ern fish­ing nets are al­most un­de­tectable to por­poises. Since these nets are com­monly used in nearshore areas, in the nat­ural range of P. pho­coena, they prob­a­bly kill large num­bers of por­poises. Var­i­ous mea­sures are being taken in the United States and other na­tions in an at­tempt to limit deaths of this kind. In ad­di­tion to deaths re­lated to fish­ing, por­poises also suf­fer from chem­i­cal and noise pol­lu­tion. (Dollinger(1988), John­ston(1999), McWilliam(1999), Nowak(1999))

Other Com­ments

One in­ter­est­ing dan­ger that the Har­bor Por­poise may be fac­ing is com­pletely nat­ural. As one group of re­searchers no­ticed, a num­ber of por­poise car­casses taken in Scot­land all had sim­i­lar punc­ture and bruise wounds, with most of them dying as a re­sult of in­ter­nal in­jury. In ad­di­tion to these in­juries some of the spec­i­mens also had teeth gouges in their flesh. After an­a­lyz­ing these bite pat­terns, the re­searchers de­ter­mined that the an­i­mal re­spon­si­ble for in­flict­ing these wounds was Tur­siops trun­ca­tus, the Bot­tlenosed Dol­phin. At least in this area of Scot­land it seems that where the two species' ranges over­lap, dol­phins are at­tack­ing and in some cases killing por­poises. It is not known for sure ex­actly what is prompt­ing this kind of be­hav­ior. (The Por­poise Page, 1998)

Con­trib­u­tors

George Ham­mond (), An­i­mal Di­ver­sity Web.

An­drew Masi (au­thor), Uni­ver­sity of Michi­gan-Ann Arbor, Phil Myers (ed­i­tor), Mu­seum of Zo­ol­ogy, Uni­ver­sity of Michi­gan-Ann Arbor.

Glossary

Arctic Ocean

the body of water between Europe, Asia, and North America which occurs mostly north of the Arctic circle.

Atlantic Ocean

the body of water between Africa, Europe, the southern ocean (above 60 degrees south latitude), and the western hemisphere. It is the second largest ocean in the world after the Pacific Ocean.

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Nearctic

living in the Nearctic biogeographic province, the northern part of the New World. This includes Greenland, the Canadian Arctic islands, and all of the North American as far south as the highlands of central Mexico.

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Pacific Ocean

body of water between the southern ocean (above 60 degrees south latitude), Australia, Asia, and the western hemisphere. This is the world's largest ocean, covering about 28% of the world's surface.

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Palearctic

living in the northern part of the Old World. In otherwords, Europe and Asia and northern Africa.

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bilateral symmetry

having body symmetry such that the animal can be divided in one plane into two mirror-image halves. Animals with bilateral symmetry have dorsal and ventral sides, as well as anterior and posterior ends. Synapomorphy of the Bilateria.

brackish water

areas with salty water, usually in coastal marshes and estuaries.

carnivore

an animal that mainly eats meat

chemical

uses smells or other chemicals to communicate

coastal

the nearshore aquatic habitats near a coast, or shoreline.

endothermic

animals that use metabolically generated heat to regulate body temperature independently of ambient temperature. Endothermy is a synapomorphy of the Mammalia, although it may have arisen in a (now extinct) synapsid ancestor; the fossil record does not distinguish these possibilities. Convergent in birds.

estuarine

an area where a freshwater river meets the ocean and tidal influences result in fluctuations in salinity.

freshwater

mainly lives in water that is not salty.

holarctic

a distribution that more or less circles the Arctic, so occurring in both the Nearctic and Palearctic biogeographic regions.

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Found in northern North America and northern Europe or Asia.

iteroparous

offspring are produced in more than one group (litters, clutches, etc.) and across multiple seasons (or other periods hospitable to reproduction). Iteroparous animals must, by definition, survive over multiple seasons (or periodic condition changes).

migratory

makes seasonal movements between breeding and wintering grounds

motile

having the capacity to move from one place to another.

natatorial

specialized for swimming

native range

the area in which the animal is naturally found, the region in which it is endemic.

pelagic

An aquatic biome consisting of the open ocean, far from land, does not include sea bottom (benthic zone).

piscivore

an animal that mainly eats fish

polar

the regions of the earth that surround the north and south poles, from the north pole to 60 degrees north and from the south pole to 60 degrees south.

saltwater or marine

mainly lives in oceans, seas, or other bodies of salt water.

seasonal breeding

breeding is confined to a particular season

sexual

reproduction that includes combining the genetic contribution of two individuals, a male and a female

social

associates with others of its species; forms social groups.

tactile

uses touch to communicate

temperate

that region of the Earth between 23.5 degrees North and 60 degrees North (between the Tropic of Cancer and the Arctic Circle) and between 23.5 degrees South and 60 degrees South (between the Tropic of Capricorn and the Antarctic Circle).

territorial

defends an area within the home range, occupied by a single animals or group of animals of the same species and held through overt defense, display, or advertisement

viviparous

reproduction in which fertilization and development take place within the female body and the developing embryo derives nourishment from the female.

Ref­er­ences

Dollinger (ed­i­tor), P. 1988. Con­ven­tion on In­ter­na­tional Trade in En­dan­gered Species of Wild Fauna and Flora; Iden­ti­fi­ca­tion Man­ual Vol I. Mam­malia. Switzer­land: Sec­re­tariat of the Con­ven­tion.

John­ston, D. 2001. "Har­bour Por­poise" (On-line). @​Phocoena.​org. Ac­cessed 11/03/04 at http://​phocoena.​org/​factsheets/​harbourporp.​html.

Nowak, R. 1999. Walker's Mam­mals of the World, 6th Ed. Vol II. Bal­ti­more: John Hop­kins Uni­ver­sity Press.

The Por­poise Page, 1998. "Har­bor Por­poise" (On-line). The Por­poise Page. Ac­cessed 11/03/04 at http://​www.​theporpoisepage.​com/​harbor.​php.