Parabuteo unicinctusHarris's hawk

Ge­o­graphic Range

Har­ris' hawks can be found in semi-open habi­tats in the south­west­ern United States and north­west­ern Mex­ico, from Baja Cal­i­for­nia to south­ern Ari­zona, New Mex­ico and Texas, ex­tend­ing south through Cen­tral and South Amer­ica to Chile and just into Patag­o­nia. (Johns­gard, 1990)

Habi­tat

Har­ris' hawks are found in var­i­ous habi­tats, from up­land desert dom­i­nated by saguaros to mesquite, palo verde, and iron­wood wood­lands in the Col­orado River val­ley. There is a pop­u­la­tion of hawks being rein­tro­duced to the Col­orado River that pre­fer to nest near water in mequite, wil­lows and cot­ton­woods. In urban areas, they are seen uti­liz­ing washes, open lots, and open desert. These hawks may be found at el­e­va­tions of 400 to 1,000 me­ters. (Driscoll, 2000; Johns­gard, 1990)

  • Range elevation
    400 to 1,000 m
    1312.34 to ft

Phys­i­cal De­scrip­tion

These are large hawks with long tails and broad wings. Har­ris's hawks range in length from 18 to 23 inches (46 to 76 cm) and have wingspans of 40 to 47 inches (100 to 120 cm). Adult plumage is uni­formly choco­late brown with dis­tinct red­dish shoul­ders, upper and un­der­wing coverts, and leg feath­ers. The tail is dark with white upper and un­der­tail coverts and a white base and ter­mi­nal band. Ju­ve­niles are sim­i­lar to adults but are less dis­tinctly col­ored and have a white belly with choco­late brown streak­ing. The tarsal feath­ers are pale with red­dish bar­ring and there is bar­ring on the tail and wings. Fe­males weigh an av­er­age of 1,047 grams, and males are smaller, weigh­ing an av­er­age of 735 grams. (Driscoll, 2000; Thomas and Gates, 1998; Wheeler and Clark, 1996)

  • Sexual Dimorphism
  • female larger
  • Range mass
    735 to 1047 g
    25.90 to 36.90 oz
  • Range length
    46 to 76 cm
    18.11 to 29.92 in
  • Range wingspan
    100 to 120 cm
    39.37 to 47.24 in

Re­pro­duc­tion

Most often, so­cial groups of Har­ris' hawks con­tain a sin­gle monog­a­mous breed­ing pair. How­ever, these hawks are known to prac­tice si­mul­ta­ne­ous polyandry, where more than one male mates with one fe­male and shares in the re­spon­si­bil­i­ties of rais­ing off­spring. Polyandry is com­monly found in areas where the habi­tat qual­ity is rich as op­posed to arid habi­tats where the chances of re­pro­duc­tive suc­cess are less, even when there are three adults hunt­ing. It is also found to be com­mon in Ari­zona where the sex ratio is sig­nif­i­cantly skewed to­wards males, in com­par­i­son with areas such as Texas, where the sex ratio is not as skewed. (Johns­gard, 1990)

Har­ris' hawks build their nests in saguaros, palo verdes and mesquite trees at an av­er­age height of 5 me­ters. In urban areas, nests can be found on cot­ton­woods, iron­woods, palm trees and elec­tri­cal tow­ers. Nests are plat­forms made of sticks, weeds, twigs, and are usu­ally lined with soft mosses, grasses and roots. Be­tween two and four eggs are laid at a time. Fe­males have the abil­ity to breed all year long and can lay two to three clutches within a year. The in­cu­ba­tion pe­riod lasts about 35 days and the males often share du­ties with the fe­male dur­ing this pe­riod. Fledg­ing oc­curs after an­other 40 days. The young birds tend to stay around the nest area for two to three months longer. (Driscoll, 2000; Johns­gard, 1990; Thomas and Gates, 1998)

  • Breeding interval
    Harris' hawks breed two to three times per year.
  • Breeding season
    Harris' hawks breed year round.
  • Range eggs per season
    3 to 15
  • Average eggs per season
    6
  • Average eggs per season
    3
    AnAge
  • Range time to hatching
    33 to 37 days
  • Range fledging age
    35 to 45 days
  • Average time to independence
    2-3 months

Both the fe­male and the male con­tribute to parental care. Har­ris' hawks prac­tice co­op­er­a­tive breed­ing, with sev­eral birds help­ing with build­ing nests, in­cu­ba­tion, feed­ing, and de­fense. This as­sis­tance in­creases nest suc­cess. There is often a trio con­sist­ing of two males and a fe­male which aid in the nest cycle. (Driscoll, 2000; Johns­gard, 1990; Thomas and Gates, 1998)

  • Parental Investment
  • pre-fertilization
    • provisioning
    • protecting
      • female
  • pre-hatching/birth
    • provisioning
      • male
      • female
    • protecting
      • male
      • female
  • pre-weaning/fledging
    • provisioning
      • male
      • female
    • protecting
      • male
      • female
  • pre-independence
    • provisioning
      • male
      • female
    • protecting
      • male
      • female

Lifes­pan/Longevity

Records on longevity are col­lected from the Bird Band­ing Lab­o­ra­tory (BBL) in Lau­rel, Mary­land. The max­i­mum longevity record for Har­ris' hawks is 14 years, 11 months. (Klimkiewicz, 2002)

Be­hav­ior

Har­ris' hawks are non-mi­gra­tory and di­ur­nal. They form com­plex so­cial groups, which aid in the nest­ing cycle. Most often these groups are trios con­sist­ing of two males and a fe­male, but groups of four or five hawks are not un­com­mon. There is a strict dom­i­nance hi­er­ar­chy within groups of Har­ris' hawks. The breed­ing, or alpha, fe­male, is dom­i­nant to all other hawks in the group. Oc­ca­sion­ally there is a sec­ond fe­male who is sub­or­di­nate to the alpha fe­male but dom­i­nant to all other males in the group. The breed­ing, or alpha, male is dom­i­nant to all other males in the group. Com­monly the group con­tains a beta male, who may at­tempt, often un­suc­cess­fully, to mate with the alpha fe­male. Fi­nally, there may be sev­eral gamma birds, which are sub­or­di­nate to the alpha and beta in­di­vid­u­als. These gamma birds may be ei­ther male or fe­male, and usu­ally they are sex­u­ally im­ma­ture in­di­vid­u­als. Often they are the ju­ve­nile off­spring of the alpha pair. All mem­bers of the group help with ob­tain­ing food, de­fend­ing the breed­ing ter­ri­tory, and pro­vid­ing nest pro­tec­tion. These groups also hunt co­op­er­a­tively. They are able to de­pend on much larger prey when hunt­ing in groups. This as­pect of group hunt­ing and food shar­ing in­creases sur­vival rates for birds as in­di­vid­u­als. (Bed­narz and David, 1988; Coul­son and Coul­son, 1995; Daw­son and Man­nan, 1991; Thomas and Gates, 1998)

  • Range territory size
    0.2 to 5.5 km^2

Home Range

Har­ris' hawks es­tab­lish and de­fend ter­ri­to­ries that range from 0.2 to 5.5 square kilo­me­ters in size. Ter­ri­tory size de­pends on the avail­abil­ity of food and other re­sources.

Com­mu­ni­ca­tion and Per­cep­tion

Like all hawks, Har­ris' hawks have keen vi­sion and hear­ing. They are known to make hiss­ing noises, give alarm calls, and prob­a­bly com­mu­ni­cate vi­su­ally as well.

Food Habits

The diet of Har­ris' hawks is ver­sa­tile and varies with prey avail­abil­ity. These hawks feed mostly on small mam­mals such as rats and mice, but also take birds and lizards. They com­monly hunt in groups of about five hawks, in­creas­ing their suc­cess rate and en­abling them to take larger prey such as cot­ton­tails and jack rab­bits. These hunt­ing groups con­sist of a breed­ing pair and other helpers, with the fe­male dom­i­nat­ing. They are fast fly­ers and once they have spot­ted their prey, they land and take turns try­ing to scare and ac­tu­ally flush the prey an­i­mal until it darts from be­neath its hid­ing place. An­other mem­ber of the hunt­ing group cap­tures the an­i­mal and as­sumes a pos­ture known as mantling, in which the hawk shields the prey with its wings to hide it from other birds. It has been sug­gested that group hunt­ing is en­cour­aged by the dense brush and thorny na­ture of their habi­tat. There is some ev­i­dence that these hawks may feed on car­rion if food avail­abil­ity is low. (Bed­narz and David, 1988; Coul­son and Coul­son, 1995; Johns­gard, 1990)

  • Primary Diet
  • carnivore
    • eats terrestrial vertebrates
  • Animal Foods
  • birds
  • mammals
  • reptiles

Pre­da­tion

Great horned owls (Bubo vir­gini­anus) pose the great­est pre­da­tion threat to this species, but coy­otes (Canis la­trans) and com­mon ravens (Corvus corax) also threaten young hawks. Fe­male Har­ris' hawks uti­lize helpers to pro­tect their nests. The helpers perch in saguaros and scan the sur­round­ings for preda­tors. They tend to be­come ex­cited and will use an alarm call when preda­tors come within their nest­ing area. Groups con­sist­ing of 2 to 5 hawks will at­tack and ha­rass any preda­tor threat­en­ing the nest. The alpha male is most likely to strike the preda­tor as the fe­male stays be­hind to pro­tect the nest. This es­tab­lish­ment of helpers greatly in­creases the de­tec­tion of preda­tors and nest suc­cess. (Daw­son and Man­nan, 1991)

Ecosys­tem Roles

Har­ris' hawks are im­por­tant preda­tors in their ecosys­tem, con­trol­ling pop­u­la­tions of many small mam­mal species. (Coul­son and Coul­son, 1995)

Eco­nomic Im­por­tance for Hu­mans: Pos­i­tive

Har­ris' hawks are of great ben­e­fit to farm­ers whose crops are de­stroyed by ro­dents. These hawks feed pri­mar­ily on small ro­dents such as mice and rats and there­fore al­le­vi­ate a lot of de­struc­tion to crops. (Coul­son and Coul­son, 1995)

  • Positive Impacts
  • controls pest population

Eco­nomic Im­por­tance for Hu­mans: Neg­a­tive

The only neg­a­tive im­pact of these hawks is their habit of con­gre­gat­ing on elec­tri­cal trans­form­ers, where they are often elec­tro­cuted. This has be­come a great cost to elec­tric com­pa­nies who are being forced to rein­su­late and, in some cases, build arms for perch­ing to re­duce the mor­tal­ity rates of hawks. (Driscoll, 2000)

Con­ser­va­tion Sta­tus

Har­ris's Hawks are not listed as threat­ened or en­dan­gered. They are in­cluded in CITES ap­pen­dix II and they are pro­tected from ha­rass­ment and il­le­gal shoot­ing by the Mi­gra­tory Bird Treaty Act. A pop­u­la­tion on the Col­orado River is thought to have been ex­tir­pated due to their de­pen­dence on a ri­par­ian com­mu­nity which was al­tered by dam con­struc­tion and dis­tur­bance from dredg­ing as well as nest de­struc­tion. Real es­tate and agri­cul­ture threaten the species in Ari­zona. Re­cent de­clines in Texas pop­u­la­tions re­sulted from the clear­ing of mesquite for agri­cul­ture and live­stock graz­ing. Habi­tat loss is the major cause of de­cline of this species as well as ex­ces­sive human dis­tur­bance. Shoot­ing can re­sult in nest fail­ure, aban­don­ment and mor­tal­ity. Elec­tro­cu­tion is re­spon­si­ble for the loss of half of the pop­u­la­tion of breed­ing hawks. It is pos­si­ble in areas such as Ari­zona for birds to live in cities where the na­tive veg­e­ta­tion is pro­tected, houses are spread apart and there is not an over­abun­dant amount of as­phalt and con­crete. (Johns­gard, 1990; Wha­ley, 1986)

Con­trib­u­tors

Al­li­son Poor (ed­i­tor), Uni­ver­sity of Michi­gan-Ann Arbor.

Mary Truglio (au­thor), Uni­ver­sity of Ari­zona, Jay Tay­lor (ed­i­tor), Uni­ver­sity of Ari­zona.

Glossary

Nearctic

living in the Nearctic biogeographic province, the northern part of the New World. This includes Greenland, the Canadian Arctic islands, and all of the North American as far south as the highlands of central Mexico.

World Map

Neotropical

living in the southern part of the New World. In other words, Central and South America.

World Map

acoustic

uses sound to communicate

bilateral symmetry

having body symmetry such that the animal can be divided in one plane into two mirror-image halves. Animals with bilateral symmetry have dorsal and ventral sides, as well as anterior and posterior ends. Synapomorphy of the Bilateria.

carnivore

an animal that mainly eats meat

chemical

uses smells or other chemicals to communicate

cooperative breeder

helpers provide assistance in raising young that are not their own

desert or dunes

in deserts low (less than 30 cm per year) and unpredictable rainfall results in landscapes dominated by plants and animals adapted to aridity. Vegetation is typically sparse, though spectacular blooms may occur following rain. Deserts can be cold or warm and daily temperates typically fluctuate. In dune areas vegetation is also sparse and conditions are dry. This is because sand does not hold water well so little is available to plants. In dunes near seas and oceans this is compounded by the influence of salt in the air and soil. Salt limits the ability of plants to take up water through their roots.

diurnal
  1. active during the day, 2. lasting for one day.
endothermic

animals that use metabolically generated heat to regulate body temperature independently of ambient temperature. Endothermy is a synapomorphy of the Mammalia, although it may have arisen in a (now extinct) synapsid ancestor; the fossil record does not distinguish these possibilities. Convergent in birds.

fertilization

union of egg and spermatozoan

iteroparous

offspring are produced in more than one group (litters, clutches, etc.) and across multiple seasons (or other periods hospitable to reproduction). Iteroparous animals must, by definition, survive over multiple seasons (or periodic condition changes).

monogamous

Having one mate at a time.

motile

having the capacity to move from one place to another.

native range

the area in which the animal is naturally found, the region in which it is endemic.

oviparous

reproduction in which eggs are released by the female; development of offspring occurs outside the mother's body.

polyandrous

Referring to a mating system in which a female mates with several males during one breeding season (compare polygynous).

riparian

Referring to something living or located adjacent to a waterbody (usually, but not always, a river or stream).

scrub forest

scrub forests develop in areas that experience dry seasons.

sedentary

remains in the same area

sexual

reproduction that includes combining the genetic contribution of two individuals, a male and a female

social

associates with others of its species; forms social groups.

suburban

living in residential areas on the outskirts of large cities or towns.

tactile

uses touch to communicate

temperate

that region of the Earth between 23.5 degrees North and 60 degrees North (between the Tropic of Cancer and the Arctic Circle) and between 23.5 degrees South and 60 degrees South (between the Tropic of Capricorn and the Antarctic Circle).

territorial

defends an area within the home range, occupied by a single animals or group of animals of the same species and held through overt defense, display, or advertisement

tropical

the region of the earth that surrounds the equator, from 23.5 degrees north to 23.5 degrees south.

tropical savanna and grassland

A terrestrial biome. Savannas are grasslands with scattered individual trees that do not form a closed canopy. Extensive savannas are found in parts of subtropical and tropical Africa and South America, and in Australia.

savanna

A grassland with scattered trees or scattered clumps of trees, a type of community intermediate between grassland and forest. See also Tropical savanna and grassland biome.

temperate grassland

A terrestrial biome found in temperate latitudes (>23.5° N or S latitude). Vegetation is made up mostly of grasses, the height and species diversity of which depend largely on the amount of moisture available. Fire and grazing are important in the long-term maintenance of grasslands.

urban

living in cities and large towns, landscapes dominated by human structures and activity.

visual

uses sight to communicate

year-round breeding

breeding takes place throughout the year

Ref­er­ences

Bed­narz, J., J. David. 1988. A Study of the Eco­log­i­cal Basis of Co­op­er­a­tive Breed­ing in the Har­ris' Hawk. Ecol­ogy, 69(4): 1176-1187.

Coul­son, J., T. Coul­son. 1995. Group Hunt­ing by Har­ris' Hawks in Texas. Jour­nal of Rap­tor Re­search, 29(4): 265-267.

Daw­son, J., R. Man­nan. 1991. Dom­i­nance Hi­er­ar­chies and Helper Con­tri­bu­tion in Har­ris' Hawks. Auk, 108(3): 649-660.

Driscoll, J. 2000. "AZGF Nongame Field Notes - Har­ris' Hawk (Parabu­teo unicinc­tus)" (On-line). Ac­cessed March 27, 2002 at http://​www.​gf.​state.​az.​us/​w_​c/​nongame_​harris_​hawk.​shtml.

Johns­gard, P. 1990. Hawks, Ea­gles, and Fal­cons of North Amer­ica. Wash­ing­ton and Lon­don: Smith­son­ian In­sti­tu­tion Press.

Klimkiewicz, M. 2002. "Longevity Records of North Amer­i­can Birds" (On-line). Ac­cessed April 18, 2002 at http://​www.​pwrc.​usgs.​gov/​bbl/​homepage/​long2890.​htm.

Thomas, A., C. Gates. 1998. "North Amer­i­can Rap­tors: Har­ris' Hawk" (On-line). Ac­cessed March 27, 2002 at http://​library.​thinkquest.​org/​18166/​harris'_hawk.​html.

Wha­ley, W. 1986. Pop­u­la­tion Ecol­ogy of the Har­ris' Hawk in Ari­zona. Jour­nal of Rap­tor Re­search, 20(1): 1-15.

Wheeler, B., W. Clark. 1996. North Amer­i­can Rap­tors. New York: Aca­d­e­mic Press.