Otus kennicottiiwestern screech owl

Ge­o­graphic Range

West­ern screech-owls (Otus ken­ni­cotti) in­habit west­ern North Amer­ica, north­west­ern Mex­ico and coastal re­gions of the Pa­cific North­west (Can­nings and An­gell, 2001). Rang­ing as far north as south­east Alaska and south­ern Canada to the Baja pen­nin­sula and into Mex­ico, west­ern screech-owls make their homes within the di­verse com­mu­ni­ties of the ri­par­ian de­cid­u­ous wood­lands of North Amer­ica (Cor­nell Lab of Or­nithol­ogy, 1999). Within the last 40 years, the range of west­ern screech-owls has ex­panded as far east as Texas (per­haps due to in­creased tree-plant­ing) and as a re­sult they are now found sym­patri­cally with their very close rel­a­tive, east­ern screech-owls (Otus asio). (Can­nings and An­gell, 2001; Cor­nell Lab of Or­nithol­ogy, 1999)

Habi­tat

West­ern screech-owls usu­ally in­habit low el­e­va­tion ri­par­ian and de­cid­u­ous oak wood­land com­m­mu­ni­ties (Camp­bell et al., 1990; Can­nings, 1997; Can­nings and An­gell, 2001). They will often in­habit stream­side groves, deserts, sub­ur­ban parks and gar­dens (Cor­nell Lab of Or­nithol­ogy, 1999). The abun­dance of O. ken­ni­cotti is pos­i­tively as­so­ci­ated with over­story cover of mesquite (prosopis) and un­der­story cover of na­tive peren­nial veg­e­ta­tion (Hardy et al., 1999).

Within the United States O. ken­ni­cot­tii oc­curs in high den­si­ties in the fol­low­ing re­gions:

Ari­zona: mesquite-ri­par­ian zones of the Sono­ran Desert. In par­tic­u­lar, west­ern screech-owls are strongly as­so­ci­ated with struc­turally di­verse up­lands of the Sono­ran Desert, con­tain­ing saguaros (Carnegiea gi­gan­tea), mesquite wood­lands and cot­ton­wood (Pop­u­lus)-wil­low (Salix) ri­par­ian areas (Hardy et al., 1999).

Texas: lower el­e­va­tions in ri­par­ian zones dom­i­nated by Ari­zona sycamore (Plan­tanus wrightii) as well as pure oak habi­tats (Gel­bach and Lev­erett, 1995; Can­nings and An­gell, 2001).

Cal­i­for­nia: west­ern screech-owls are strongly as­so­ci­ated with areas with fan palms (Wash­ing­to­nia fil­if­era) and oak wood­lands (Feusier, 1989; Noble, 1990; Lehman, 1994; Can­nings and An­gell, 2001).

Wash­ing­ton and Alaska: west­ern screech-owls are found in ri­par­ian habi­tats and the mixed forests of coastal re­gions con­sist­ing of big-leaf maple (Acer macro­phyl­lum), red alder (Alnus rubra), Dou­glas fir (Psu­dot­suga men­ziesii), west­ern hem­lock (Tsuga het­ero­phylla)and west­ern red cedar (Thyja pli­cata) (Kitchin, 1949; Can­nings and An­gell, 2001).

Col­orado: west­ern screech-owls are pri­mar­ily found in rural de­vel­oped areas and areas with broad-leaved cot­ton­woods along river basins. They are also as­so­ci­ated with farm­yards con­tain­ing ma­ture, un­pruned cot­ton­wood trees and shrub­bery. Through­out their range they are found in urban and sub­ur­ban parks as well as res­i­den­tial areas (Levad, 1998; Can­nings and An­gell, 2001). (Camp­bell, et al., 1990; Can­nings and An­gell, 2001; Cor­nell Lab of Or­nithol­ogy, 1999; Feusier, 1989; Gehlbach and Lev­erett, 1995; Hardy, et al., 1999; Kitchin, 1949; Lehman, 1994; Levad, 1998; Noble, 1990)

Phys­i­cal De­scrip­tion

Otus ken­ni­cot­tii is a small owl with feath­ered "ear tufts". Adults are 19 to 25.5 cm in length; and weigh 170.1 g on av­er­age. Male wingspans are 168.4 mm, fe­male wingspans are 174.5 mm. Sexes are alike in plumage char­ac­ter­is­tics. Plumage of west­ern screech-owls is gen­er­ally monomor­phhic in a given area; brown or gray-brown in the north­west and gray in south­ern deserts. Some pop­u­la­tions in the coastal re­gions of the Pa­cific North­west are more vari­able in color, often dis­play­ing red­dish-brown morphs (Can­nings and An­gell, 2001).

The face of west­ern screech-owls is pale with a dark lat­eral bor­der, their un­der­parts are streaked and barred. They have yel­low eyes and dark bills. Their feet and toes are feath­ered in north­ern pop­u­la­tions but bris­tled in south­ern deserts (Can­nings and An­gell, 2001). (Can­nings and An­gell, 2001)

  • Sexual Dimorphism
  • sexes alike
  • female larger
  • Range mass
    163.5 to 183 g
    5.76 to 6.45 oz
  • Average mass
    170.1 g
    5.99 oz
  • Range length
    19 to 25.5 cm
    7.48 to 10.04 in
  • Range wingspan
    168.4 to 174.5 mm
    6.63 to 6.87 in

De­vel­op­ment

De­vel­op­ment of young at age-

1 week: eyes open and egg tooth ab­sent.

1-2 weeks: young bill snap in re­sponse to dis­tur­bance.

3 weeks: more ag­gres­sive, hiss­ing and sway­ing with wings out­streched.

Reach fledg­ling state within ap­prox­i­mately 35 days. For the first 5 weeks fol­low­ing fledg­ing, ju­ve­nilles will re­main in close as­so­ci­a­tion with thier par­ents and most prob­a­bly begin breed­ing when 1 year old (Sum­ner 1929, Can­nings and An­gell 2001).

Re­pro­duc­tion

Otus ken­ni­cot­tii is usu­ally mono­go­mous, al­though polygamy oc­curs oc­ca­sion­ally.

Be­fore the breed­ing sea­son, male O. ken­ni­cot­tii de­fend an area con­tain­ing sev­eral nest cav­i­ties. Pair for­ma­tion be­gins dur­ing the months of Jan­u­ary and Feb­ru­ary through­out the range. Al­lo­preen­ing is an im­por­tant part of con­tin­u­ous pair-bond­ing be­hav­ior and oc­curs at all times of the year. Courtship-feed­ing is com­mon. The male will pre­sent food to the fe­male and per­form elab­o­rate courtship dis­plays in­volv­ing bow­ing, bill snap­ping, and hop­ping. (Cor­nell Lab of Or­nithol­ogy, 1999)

Through­out its range, O. ken­ni­cot­tii will nest in tree cav­i­ties, most com­monly those ex­ca­vated by other species. Ex­am­ples in­clude: fox squir­rels, north­ern flick­ers, gilded flick­ers, pileated wood­peck­ers and Eu­ro­pean star­lings (Camp­bell et al., 1990; Can­nings and An­gell, 2001; Cor­nell Lab of Or­nithol­ogy, 1999). They also in­habit nat­ural tree cavites and nest boxes.

Eggs are laid dur­ing March and April. They are white and oval to broadly elip­ti­cal. Fe­males lay 2 to 7 eggs per clutch (3 to 5 on av­er­age). In­cu­ba­tion lasts 26 to 34 days and the chicks fledge in 27 to 35 days.

West­ern screech-owls prob­a­bly begin breed­ing when they are 1 year old, adults at­tempt to nest every year (Can­nings and An­gell, 2001). (Camp­bell, et al., 1990; Can­nings and An­gell, 2001; Cor­nell Lab of Or­nithol­ogy, 1999)

  • Breeding interval
    Western screech owls breed once yearly.
  • Breeding season
    Eggs are laid in March or April.
  • Range eggs per season
    2 to 7
  • Average eggs per season
    3
  • Range time to hatching
    26 to 34 days
  • Range fledging age
    29 to 35 days
  • Average age at sexual or reproductive maturity (female)
    1 years
  • Average age at sexual or reproductive maturity (male)
    1 years

West­ern screech-owls add no new ma­te­r­ial to the nest site, rather they use what­ever sub­strate is al­ready pre­sent. They pre­fer to nest in tree cav­i­ties or nest boxes. Cot­ton­wood seems to be fa­vored wher­ever avail­able, most likely be­cause of its ten­dency to form large nat­ural cav­i­ties (Can­nings and An­gell, 2001).

Al­tri­cial nestlings are cov­ered in white down when they hatch. Their eyes are closed and they have an egg tooth. Males will feed fe­males through­out egg-lay­ing, in­cu­ba­tion and brood­ing pe­ri­ods until the young are about 3 weeks old (Can­nings and An­gell, 2001). Both males and fe­males pro­vide food for the nestlings (Cor­nell Lab of Or­nithol­ogy, 1999; Ehrlich et al., 1988). (Can­nings and An­gell, 2001; Cor­nell Lab of Or­nithol­ogy, 1999; Ehrlich, et al., 1988)

  • Parental Investment
  • no parental involvement
  • altricial
  • pre-fertilization
  • pre-hatching/birth
    • protecting
      • female
  • pre-weaning/fledging
    • provisioning
      • male
      • female
    • protecting
      • female

Lifes­pan/Longevity

West­ern screech-owls usu­ally live from 1 to 8 years in the wild. The longest recorded lifes­pan for a west­ern screech-owl in the wild is 13 years, in cap­tiv­ity the longest recorded lifes­pan is 19 years.

  • Range lifespan
    Status: wild
    13 (high) years
  • Range lifespan
    Status: captivity
    19 (high) years
  • Typical lifespan
    Status: wild
    1 to 8 years

Be­hav­ior

The wings of Otus ken­ni­cot­tii have a low as­pect ratio, which is typ­i­cal of sit-and-wait preda­tors that do not fly long dis­tances (John­son, 1997; Can­nings and An­gell, 2001). Young are very ca­pa­ble climbers and adults are able to ef­fec­tively walk into thick­ets for roost­ing (Can­nings and An­gell, 2001).

Otus ken­ni­cot­tii has been ob­served to bathe oc­ca­sion­ally. Males roost in close prox­im­ity to the nest dur­ing breed­ing sea­son, this re­duces the fly­ing dis­tance to the nest through­out the nestling pe­riod. Males are joined by the fe­males dur­ing the last weeks of the nestling pe­riod and they sys­tem­at­i­cally po­si­tion their roosts op­po­site the nest cav­ity en­trance to pro­vide in­creased pro­tec­tion from crows, jays, and other di­ur­nal preda­tors (Can­nings and An­gell; 2001).

Ter­ri­to­ries are de­fended year-round, how­ever, re­sponses to calls by con­specifics de­cline dur­ing the in­cu­ba­tion and nestling pe­ri­ods. West­ern screech-owls react agres­sively to calls of spot­ted owls (Strix oc­ci­den­talis) and great horned owls (Bubo vir­gini­anus) (Can­nings and An­gell; 2001).

Through­out the year, west­ern screech-owls live in pairs. How­ever, they can be found in small fam­ily groups be­fore the ju­ve­nilles dis­perse (Can­nings and An­gell; 2001). (Can­nings and An­gell, 2001; John­son, 1997)

Home Range

We do not have in for­ma­tion on home range for this species at this time.

Com­mu­ni­ca­tion and Per­cep­tion

The song of Otus ken­ni­cot­tii con­sists of 5 to 15 hol­low whis­tle hoots of a sin­gle pitch that ac­cel­er­ate to­wards the end in a "bounc­ing ball" se­quence (Can­nings and An­gell, 2001). (Can­nings and An­gell, 2001)

Food Habits

West­ern screech-owls are "sit-and-wait" preda­tors. They leave their roosts to for­age just be­fore sun­set. Their hunt­ing tech­nique may pre­dis­pose the north­ern pop­u­la­tions to pri­mar­ily hunt mice. When for­ag­ing near bod­ies of water, O. ken­ni­cot­tii will perch and wait until cray­fish emerge in the shal­lows, then it will fly down and grab one by dip­ping only its legs into the water. It will also hawk for fly­ing in­sects and glean arthro­pods from fo­lage, but it prefers to catch small, ter­res­trial mam­mals (Hay­ward and Gar­ton, 1988; Can­nings and An­gell, 2001).

Song­birds are a fre­quent meal for O. ken­ni­cot­tii and as a re­sult the owls are often mobbed in re­sponse to their threats. Fre­quent mob­bers are those song­birds that are most often preyed upon (for ex­am­ple, north­ern car­di­nals and white-throated spar­rows). Mob­bing by song­birds oc­curs most fre­quently dur­ing the spring and early sum­mer nest­ing pe­riod. Most mob­bers are per­ma­nent res­i­dents that are al­ways part of the avian prey com­mu­nity. Birds use sea­sonal song, age-re­alted plumage and nest-area cues of west­ern screech-owls to help as­sess the dan­ger of pre­da­tion (Gehlbach and Lev­erett, 1995).

West­ern screech-owls com­monly eat: small mam­mals, birds, an­nelid worms, in­sects, cray­fish and fish. (Can­nings and An­gell, 2001; Gehlbach and Lev­erett, 1995; Hay­ward and Gar­ton, 1988)

  • Animal Foods
  • birds
  • mammals
  • fish
  • insects
  • terrestrial non-insect arthropods
  • terrestrial worms

Pre­da­tion

West­ern screech-owls typ­i­cally react to the pres­ence of preda­tors with bill snaps and bark calls.

Known preda­tors in­clude: spot­ted owls (Strix oc­ci­den­talis), great horned owls (Bubo vir­gini­anus), barred owls (Strix varia), rac­coons (Pro­cyon lotor), go­pher snakes (Pituophis melanoleu­cus) and crows and jays (fam­ily Corvi­dae). (Can­nings and An­gell, 2001)

Ecosys­tem Roles

Colonies of tree ants (genus Cre­mato­gaster) will often oc­cupy the nest sites of Otus ken­ni­cot­tii and ap­pa­rantly pro­tect the nest from dis­tur­bance by bit­ing or sting­ing po­ten­tial preda­tors. This re­la­tion­ship seems to be a unique sym­bio­sis in the genus Otus, how­ever, it is in­fre­quent (Mc­Cal­lum et al., 1995).

The abun­dance of west­ern screech-owls is pos­i­tively as­so­ci­ated with the abun­dance of elf owls (Mi­cra­thene whit­neyi) in the Sono­ran Desert (Hardy et al., 1999). (Hardy, et al., 1999; Mc­Cal­lum, et al., 1995)

Mu­tu­al­ist Species

Eco­nomic Im­por­tance for Hu­mans: Pos­i­tive

West­ern screech-owls can live in sub­ur­ban habi­tats where nest and root sites are avail­able. Pairs nest­ing in these sub­ur­ban habi­tats are often tol­er­ant of hu­mans close to their nests , this al­lows bird­ers to visit their nest sites (Can­nings and An­gell, 2001). In ad­di­tion, Otus ken­ni­cot­tii (pre­vi­ously thought to be con­spe­cific with Otus asio), chal­lenges re­searchers to dis­cover more about this mag­nif­i­cent bird and hope­fully urge con­ser­va­tion­ists to adopt an ac­tive role in habi­tat preser­va­tion. (Can­nings and An­gell, 2001)

  • Positive Impacts
  • ecotourism
  • research and education

Eco­nomic Im­por­tance for Hu­mans: Neg­a­tive

There are no known ad­verse af­fects of west­ern screech-owls on hu­mans.

Con­ser­va­tion Sta­tus

West­ern screech-owls are closely linked to the ri­par­ian habi­tats of its range; these areas are often the first habi­tat in any given area to suf­fer the ef­fects of urban de­vel­op­ment. Al­though pairs nest­ing in sub­ur­ban areas are gen­er­ally tol­er­ant of hu­mans close to their nests, they may be sen­si­tive to local dis­tur­bance at nest sites from fre­quent vis­its by bird­ers (Hardy et al., 1999; Can­nings and An­gell, 2001). Breed­ing pop­u­la­tions are con­tin­u­ally threa­t­end by rapid ur­ban­iza­tion and degra­da­tion of habi­tat, and face pos­si­ble com­pe­ti­tion from ex­otic species such as Eu­ro­pean star­lings (Stur­nus vul­garis) (Hardy et al., 1999).

Cur­rently there are no of­fi­cial data on the trends of pop­u­la­tion den­si­ties for west­ern screech-owls. How­ever, their pop­u­la­tions are prob­a­bly de­clin­ing slowly as habi­tat is lost (Can­nings and An­gell, 2001).

West­ern screech-owls are pro­tected by the US MBTA and are listed under Ap­pen­dix II by CITES, but are not listed on the US Fed­eral List or the IUCN Red List. (Can­nings and An­gell, 2001; Hardy, et al., 1999)

Other Com­ments

West­ern screech-owls (Otus ken­ni­cot­tii), long con­sid­ered con­spe­cific with east­ern screech-owls (Otus asio), were fi­nally rec­og­nized as a seper­ate species in 1983. Their re-clas­si­fi­ca­tion was based on sig­nifi­gant dif­fer­ences in vo­cal­iza­tion and be­hav­ior be­tween the two species (Can­nings and An­gell, 2001).

De­spite its abun­dance, lit­tle is known about Otus ken­ni­cot­tii, in­clud­ing basic pa­ra­me­ters such as sur­vival rates and breed­ing suc­cess. Pri­or­i­ties for fu­ture re­search in­clude vo­cal­iza­tion stud­ies that could focus on the ge­o­graphic vari­a­tion in calls, and ge­netic analy­sis of var­i­ous sub­species for tax­o­nomic pur­poses (Can­nings and An­gell, 2001). (Can­nings and An­gell, 2001)

Con­trib­u­tors

Alaine Cam­field (ed­i­tor), An­i­mal Di­ver­sity Web.

Erin Quist (au­thor), Uni­ver­sity of Ari­zona, Todd McWhorter (ed­i­tor), Uni­ver­sity of Ari­zona.

Glossary

Nearctic

living in the Nearctic biogeographic province, the northern part of the New World. This includes Greenland, the Canadian Arctic islands, and all of the North American as far south as the highlands of central Mexico.

World Map

acoustic

uses sound to communicate

agricultural

living in landscapes dominated by human agriculture.

altricial

young are born in a relatively underdeveloped state; they are unable to feed or care for themselves or locomote independently for a period of time after birth/hatching. In birds, naked and helpless after hatching.

bilateral symmetry

having body symmetry such that the animal can be divided in one plane into two mirror-image halves. Animals with bilateral symmetry have dorsal and ventral sides, as well as anterior and posterior ends. Synapomorphy of the Bilateria.

carnivore

an animal that mainly eats meat

chemical

uses smells or other chemicals to communicate

coastal

the nearshore aquatic habitats near a coast, or shoreline.

desert or dunes

in deserts low (less than 30 cm per year) and unpredictable rainfall results in landscapes dominated by plants and animals adapted to aridity. Vegetation is typically sparse, though spectacular blooms may occur following rain. Deserts can be cold or warm and daily temperates typically fluctuate. In dune areas vegetation is also sparse and conditions are dry. This is because sand does not hold water well so little is available to plants. In dunes near seas and oceans this is compounded by the influence of salt in the air and soil. Salt limits the ability of plants to take up water through their roots.

ecotourism

humans benefit economically by promoting tourism that focuses on the appreciation of natural areas or animals. Ecotourism implies that there are existing programs that profit from the appreciation of natural areas or animals.

endothermic

animals that use metabolically generated heat to regulate body temperature independently of ambient temperature. Endothermy is a synapomorphy of the Mammalia, although it may have arisen in a (now extinct) synapsid ancestor; the fossil record does not distinguish these possibilities. Convergent in birds.

fertilization

union of egg and spermatozoan

forest

forest biomes are dominated by trees, otherwise forest biomes can vary widely in amount of precipitation and seasonality.

insectivore

An animal that eats mainly insects or spiders.

internal fertilization

fertilization takes place within the female's body

iteroparous

offspring are produced in more than one group (litters, clutches, etc.) and across multiple seasons (or other periods hospitable to reproduction). Iteroparous animals must, by definition, survive over multiple seasons (or periodic condition changes).

monogamous

Having one mate at a time.

motile

having the capacity to move from one place to another.

native range

the area in which the animal is naturally found, the region in which it is endemic.

nocturnal

active during the night

oviparous

reproduction in which eggs are released by the female; development of offspring occurs outside the mother's body.

piscivore

an animal that mainly eats fish

riparian

Referring to something living or located adjacent to a waterbody (usually, but not always, a river or stream).

seasonal breeding

breeding is confined to a particular season

sexual

reproduction that includes combining the genetic contribution of two individuals, a male and a female

suburban

living in residential areas on the outskirts of large cities or towns.

tactile

uses touch to communicate

temperate

that region of the Earth between 23.5 degrees North and 60 degrees North (between the Tropic of Cancer and the Arctic Circle) and between 23.5 degrees South and 60 degrees South (between the Tropic of Capricorn and the Antarctic Circle).

terrestrial

Living on the ground.

territorial

defends an area within the home range, occupied by a single animals or group of animals of the same species and held through overt defense, display, or advertisement

visual

uses sight to communicate

Ref­er­ences

Camp­bell, R., N. Dawe, I. Mc­Tag­gert-Cowan, J. Cooper, G. Kaiser. 1990. The birds of British Co­lum­bia. Vol 2: di­ur­nal birds of prey through wood­peck­ers. Co­lum­bia Mu­seum, Vic­to­ria: R. Br.

Can­nings, R. 1997. A sur­vey of the West­ern-Screech Owl (Otus ken­ni­cot­tii mac­far­leanei) in the in­te­rior of British Co­lum­bia. Lands and Parks, Vic­to­ria, British Co­lum­bia: Un­publ. rep., Min­istry of En­vi­ron­ment.

Can­nings, R., T. An­gell. 2001. West­ern Screech-Owl (Otus Ken­ni­cot­tii). In the Birds of North Amer­ica, No. 597. Philadel­phia, PA: (A. Poole and F. Gill, eds.). The Birds of North Amer­ica, Inc.

Cor­nell Lab of Or­nithol­ogy, 1999. "The Bird­house Net­work-East­ern Screech-Owl" (On-line). Ac­cessed 01/23/04 at http://​birds.​cornell.​edu/​birdhouse/​speciesaccounts/​EASTSCREECHOWL.​htm.

Ehrlich, P., D. Dobkin, D. Wheye. 1988. The Birder's Hand­book: A field guide to the nat­ural his­tory of North Amer­i­can birds. New York: Simon & Schus­ter Inc.

Feusier, S. 1989. Dis­tri­b­u­tion and be­hav­ior of West­ern Screech-Owls (Otus ken­ni­cot­tii) of the Starr Ranch Audubon Sanc­tu­ary, Or­ange Co., Cal­i­for­nia. Hum­boldt State Uni­ver­sity, CA: M.S. the­sis.

Gehlbach, F., J. Lev­erett. 1995. Mob­bing of east­ern screech-owls: preda­tory cues, risk to mob­bers and de­gree of threat. The Con­dor, 97(3): 831-834.

Hardy, P., M. Mar­ri­son, R. Barry. 1999. Abun­dance and habi­tat as­so­ci­a­tions of Elf Owls and West­ern Screech-Owls in the Sono­ran Desert. The South­west­ern Nat­u­ral­ist, 44(3): 311-323.

Hay­ward, G., E. Gar­ton. 1988. Re­source par­ti­tion­ing among for­est owls in the River of No Re­turn Wilder­ness, Idaho. Oe­co­log­ica, 75: 253-265.

John­son, D. 1997. Wing load­ing in 15 species of North Amer­i­can owls. Pp. 553-561 in Bi­ol­ogy and con­ser­va­tion of owls of the North­ern Hemi­sphere: 2nd In­ter­na­tional Sym­po­sium; 1997 Feb­ru­ary 5-9; Win­nipeg, MB. Stn. Gen. Tech. Rep. NC-190, St. Paul, MN: (J.R. dun­can, D.H. John­son and T.H. Nichols, eds.) USDA For­est Ser­vice, North Cen­tral Res.

Kitchin, E. 1949. Birds of the Olympic Penin­sula. Port An­ge­les, WA: Olympic Sta­tion­ers.

Lehman, P. 1994. The birds of Santa Bar­bara Co., Cal­i­for­nia. Uni­ver­sity of Cal­i­for­nia, Santa Bar­bara: Ver­te­brate Mu­seum.

Levad, R. 1998. West­ern Screech-Owl. Pp 212-213 in Col­orado breed­ing bird atlas (H.E. Kingery, ed.). , Den­ver: Col­orado Bird Atlas Part­ner­ship and Col­orado Di­vi­sion of Wildlife.

Mc­Cal­lum, D., F. Gehlbach, S. Webb. 1995. Life his­tory and ecol­ogy of Flam­mu­lated Owls in a mar­ginal New Mex­ico pop­u­la­tion. Wil­son Bul­letin, 107(3): 530-537.

Noble, P. 1990. Dis­tri­b­u­tion and den­sity of owls at Monte Bello Open Space Pre­serve, Santa Clara Co., Cal­i­for­nia. West­ern Birds, 21: 11-16.

Sum­ner, E. 1929. Com­par­a­tive stud­ies on the growth of young rap­tors. The Con­dor, 31: 85-111.