Lepus americanussnowshoe hare

Ge­o­graphic Range

Snow­shoe hares are found through­out Canada and in the north­ern­most United States. The range ex­tends south along the Sier­ras, Rock­ies, and Ap­palachian moun­tain ranges. (Kurta, 1995; Wil­son and Ruff, 1999)

Habi­tat

Snow­shoe hares are most often found in open fields, fence rows, swamps, river­side thick­ets, cedar bogs and conif­er­ous low­lands. (Kurta, 1995; Wil­son and Ruff, 1999)

Phys­i­cal De­scrip­tion

Snow­shoe hares range in length from 413 to 518 mm, of which 39 to 52 mm are tail. The hind foot, long and broad, mea­sures 117 to 147 mm in length. The ears are 62 to 70 mm from notch to tip. Snow­shoe hares usu­ally weigh be­tween 1.43 and 1.55 kg. Males are slightly smaller than fe­males, as is typ­i­cal for lep­orids. In the sum­mer, the coat is a griz­zled rusty or gray­ish brown, with a black­ish mid­dor­sal line, buffy flanks and a white belly. The face and legs are cin­na­mon brown. The ears are brown­ish with black tips and white or creamy bor­ders. Dur­ing the win­ter, the fur is al­most en­tirely white, ex­cept for black eye­lids and the black­ened tips on the ears. The soles of the feet are densely furred, with stiff hairs (form­ing the snow­shoe) on the hind feet. (Kurta, 1995; Wil­son and Ruff, 1999)

  • Sexual Dimorphism
  • female larger
  • Range mass
    1.43 to 1.55 kg
    3.15 to 3.41 lb
  • Range length
    413 to 518 mm
    16.26 to 20.39 in
  • Average basal metabolic rate
    6.708 W
    AnAge

Re­pro­duc­tion

Groups of males con­gre­gate around es­trus fe­males, fol­low­ing the fe­males as they move about their home ranges. Mat­ing is polyg­y­nan­drous (both males and fe­males have mul­ti­ple mates). (Kurta, 1995; Wil­son and Ruff, 1999)

Breed­ing sea­son for snow­shoe hares runs from mid-March through Au­gust, when the testes of the male begin to regress. Ges­ta­tion lasts 36 days. When par­tu­ri­tion ap­proaches, fe­male hares be­come highly ag­gres­sive and in­tol­er­ant of males. They re­tire to a birthing area, where they have pre­pared an area of packed down grasses. Fe­males give birth to lit­ters of up to 8 young, al­though the av­er­age lit­ter size is usu­ally two to four young. Lit­ters born late in the sea­son tend to be larger than lit­ters born in the spring. Fe­males are polye­strous and may have up to four lit­ters a year, de­pend­ing on en­vi­ro­men­tal con­di­tions. Males and fe­males be­come ma­ture within a year of their birth. (Kurta, 1995; Wil­son and Ruff, 1999)

  • Breeding interval
    Female snowshoe hares may give birth every month during the breeding season.
  • Breeding season
    Breeding season for snowshoe hares runs from mid-March through August.
  • Range number of offspring
    1 to 7
  • Average number of offspring
    2.82
  • Average number of offspring
    3
    AnAge
  • Range gestation period
    36 to 40 days
  • Average gestation period
    37.2 days
  • Range weaning age
    14 to 28 days
  • Range age at sexual or reproductive maturity (female)
    1 (high) years
  • Range age at sexual or reproductive maturity (male)
    1 (high) years

Young snow­shoe hares are pre­co­cial. They are born fully furred and able to lo­co­mote. The young hide in sep­a­rate lo­ca­tions dur­ing the day, only com­ing to­gether for 5 to 10 min­utes at a time to nurse. The fe­male alone cares for them until they are weaned and dis­perse, about four weeks after they are born. (Kurta, 1995)

  • Parental Investment
  • precocial
  • pre-fertilization
    • provisioning
    • protecting
      • female
  • pre-hatching/birth
    • provisioning
      • female
    • protecting
      • female
  • pre-weaning/fledging
    • provisioning
      • female
  • pre-independence
    • provisioning
      • female

Lifes­pan/Longevity

In the wild as much as 85% of snow­shoe hares do not live longer than one year. In­di­vid­u­als may live up to 5 years in the wild. (Carey and Judge, 2002; Kurta, 1995)

Be­hav­ior

Snow­shoe hares are typ­i­cally soli­tary, but they often live at high den­si­ties, and in­di­vid­u­als share over­lap­ping home ranges. They are ac­tive at low light lev­els and so are most often seen out and about at dawn, dusk, and dur­ing the night. They are also ac­tive on cloudy days.

Dur­ing the day­light hours, hares spend a great deal of time groom­ing, and they take fit­ful naps. Most ac­tiv­ity is re­stricted to path­ways, tram­pled down "roads" in the veg­e­ta­tion that the hares know very thor­oughly.

Hares like to take dust baths. These help to re­move ec­topar­a­sites from the hares' fur.

Snow­shoe hares are also ac­com­plished swim­mers. They oc­ca­sion­ally swim across small lakes and rivers, and they have been seen en­ter­ing the water in order to avoid preda­tors. (Kurta, 1995; Wil­son and Ruff, 1999)

  • Range territory size
    0.03 to 0.07 km^2

Home Range

Dur­ing its ac­tive pe­riod, a hare may cover up to 0.02 square kilo­me­ters of its 0.03 to 0.07 square kilo­me­ter home range.

Com­mu­ni­ca­tion and Per­cep­tion

Snow­shoe hares have acute hear­ing, which pre­sum­ably helps them to iden­tify ap­proach­ing preda­tors. They are not par­tic­u­larly vocal an­i­mals, but may make loud squeal­ing sounds when cap­tured. When en­gag­ing in ag­gres­sive ac­tivites, these an­i­mals may hiss and snort. Most com­mu­ni­ca­tion be­tween hares in­volves thump­ing the hind feet against the ground. (Kurta, 1995)

Food Habits

The diet of snow­shoe hares is vari­able. They browse on green grasses, forbs, blue­grass, brome, vetches, asters, jew­el­weed, wild straw­berry, pussy-toes, dan­de­lions, clovers, daisies and horse­tails. The new growth of trem­bling aspen, birches and wil­lows is also eaten. Dur­ing the win­ter, snow­shoe hares for­age on buds, twigs, bark, and ever­greens. They have been known to can­ni­bal­ize the re­mains of dead con­specifics in win­ter months. At all times, it is im­por­tant for hares to reingest cer­tain feces. Be­cause much of the di­ges­tion of food oc­curs in their hindguts, in order to ex­tract all of the avail­able nu­tri­ents from their food, they must cycle it through their di­ges­tive sys­tem a sec­ond time. (Kurta, 1995; Wil­son and Ruff, 1999)

  • Plant Foods
  • leaves
  • wood, bark, or stems
  • flowers
  • Other Foods
  • dung

Pre­da­tion

Snow­shoe hares are ex­perts at es­cap­ing preda­tors. Young hares often "freeze" in their tracks when they are alerted to the pres­ence of a preda­tor. Pre­sum­ably, they are at­tempt­ing to es­cape no­tice by being cryp­tic. Given the hare's back­ground-match­ing col­oration, this strat­egy is quite ef­fec­tive. Older hares are more likely to es­cape preda­tors by flee­ing. At top speed, a snow­shoe hare can travel up to 27 mile per hour. An adult hare can cover up to 10 feet in a sin­gle bound. In ad­di­tion to high speeds, hares em­ploy skill­ful changes in di­rec­tion and ver­ti­cal leaps, which may cause a preda­tor to mis­judge the exact po­si­tion of the an­i­mal from one mo­ment to the next.

Im­por­tant preda­tors of snow­shoe hares in­clude gray foxes, red foxes, coy­otes, wolves, lynx, bob­cats and mink. (Kurta, 1995; Wil­son and Ruff, 1999)

  • Anti-predator Adaptations
  • cryptic

Ecosys­tem Roles

Snow­shoe hares are im­por­tant prey an­i­mals in their ecosys­tem. (Kurta, 1995; Wil­son and Ruff, 1999)

Eco­nomic Im­por­tance for Hu­mans: Pos­i­tive

Snow­shoe hares are uti­lized widely as a source of wild meat. In ad­di­tion to this, they are an im­por­tant prey species for many preda­tors whose furs are highly val­ued. (Kurta, 1995)

  • Positive Impacts
  • food

Eco­nomic Im­por­tance for Hu­mans: Neg­a­tive

Hares may dam­age trees, es­pe­cially dur­ing pe­ri­ods of high pop­u­la­tion den­sity. (Kurta, 1995)

Con­ser­va­tion Sta­tus

Snow­shoes hares are com­mon through­out their range. Their rapid re­pro­duc­tion makes it un­likely that they will be­come a major con­cern for con­ser­va­tion­ists. (Kurta, 1995; Wil­son and Ruff, 1999)

Other Com­ments

Snow­shoe hares have been widely stud­ied. One of the more in­ter­est­ing things known about hares are the dra­matic pop­u­la­tion cy­cles that they un­dergo. Pop­u­la­tion den­si­ties can vary from 1 to 10,000 hares per square mile. The am­pli­tude of the pop­u­la­tion fluc­tu­a­tions varies across the ge­o­graphic range. It is great­est in north­west­ern Canada, and least in the rocky Moun­tain re­gion of the United States, per­haps be­cause there is more bi­o­log­i­cal di­ver­sity in more southerly re­gions. The lack of di­ver­sity in the North­west­ern por­tion of the hare's range means that there are fewer links in the food chain, and there­fore fewer species to buffer ei­ther dra­matic pop­u­la­tion in­creases or de­creases. Dis­ease may play a part in pop­u­la­tion fluc­tu­a­tion. Pneu­mono­coc­cus, ring­worm, and sal­mo­nella have all been as­so­ci­ated with pop­u­la­tion crashes.

Snow­shoe hares are also fa­mous for their sea­sonal molts. In the sum­mer, the coat of the hare is red­dish brown or gray, but dur­ing the win­ter, the coat is snowy white. The molt usu­ally takes about 72 days to reach com­ple­tion, and it seems to be reg­u­lated by daylength. In­ter­est­ingly, there seem to be two en­tirely dif­fer­ent sets of hair fol­li­cles, which give rise to white and brown hairs, re­spec­tively. (Kurta, 1995; Wil­son and Ruff, 1999)

Con­trib­u­tors

Al­li­son Poor (ed­i­tor), Uni­ver­sity of Michi­gan-Ann Arbor.

Nancy Shef­ferly (au­thor), An­i­mal Di­ver­sity Web.

Glossary

Nearctic

living in the Nearctic biogeographic province, the northern part of the New World. This includes Greenland, the Canadian Arctic islands, and all of the North American as far south as the highlands of central Mexico.

World Map

acoustic

uses sound to communicate

agricultural

living in landscapes dominated by human agriculture.

bilateral symmetry

having body symmetry such that the animal can be divided in one plane into two mirror-image halves. Animals with bilateral symmetry have dorsal and ventral sides, as well as anterior and posterior ends. Synapomorphy of the Bilateria.

bog

a wetland area rich in accumulated plant material and with acidic soils surrounding a body of open water. Bogs have a flora dominated by sedges, heaths, and sphagnum.

carrion

flesh of dead animals.

chemical

uses smells or other chemicals to communicate

crepuscular

active at dawn and dusk

cryptic

having markings, coloration, shapes, or other features that cause an animal to be camouflaged in its natural environment; being difficult to see or otherwise detect.

endothermic

animals that use metabolically generated heat to regulate body temperature independently of ambient temperature. Endothermy is a synapomorphy of the Mammalia, although it may have arisen in a (now extinct) synapsid ancestor; the fossil record does not distinguish these possibilities. Convergent in birds.

fertilization

union of egg and spermatozoan

folivore

an animal that mainly eats leaves.

food

A substance that provides both nutrients and energy to a living thing.

forest

forest biomes are dominated by trees, otherwise forest biomes can vary widely in amount of precipitation and seasonality.

herbivore

An animal that eats mainly plants or parts of plants.

iteroparous

offspring are produced in more than one group (litters, clutches, etc.) and across multiple seasons (or other periods hospitable to reproduction). Iteroparous animals must, by definition, survive over multiple seasons (or periodic condition changes).

motile

having the capacity to move from one place to another.

native range

the area in which the animal is naturally found, the region in which it is endemic.

nocturnal

active during the night

polygynandrous

the kind of polygamy in which a female pairs with several males, each of which also pairs with several different females.

saltatorial

specialized for leaping or bounding locomotion; jumps or hops.

seasonal breeding

breeding is confined to a particular season

sedentary

remains in the same area

sexual

reproduction that includes combining the genetic contribution of two individuals, a male and a female

solitary

lives alone

swamp

a wetland area that may be permanently or intermittently covered in water, often dominated by woody vegetation.

tactile

uses touch to communicate

taiga

Coniferous or boreal forest, located in a band across northern North America, Europe, and Asia. This terrestrial biome also occurs at high elevations. Long, cold winters and short, wet summers. Few species of trees are present; these are primarily conifers that grow in dense stands with little undergrowth. Some deciduous trees also may be present.

temperate

that region of the Earth between 23.5 degrees North and 60 degrees North (between the Tropic of Cancer and the Arctic Circle) and between 23.5 degrees South and 60 degrees South (between the Tropic of Capricorn and the Antarctic Circle).

terrestrial

Living on the ground.

territorial

defends an area within the home range, occupied by a single animals or group of animals of the same species and held through overt defense, display, or advertisement

tundra

A terrestrial biome with low, shrubby or mat-like vegetation found at extremely high latitudes or elevations, near the limit of plant growth. Soils usually subject to permafrost. Plant diversity is typically low and the growing season is short.

visual

uses sight to communicate

viviparous

reproduction in which fertilization and development take place within the female body and the developing embryo derives nourishment from the female.

young precocial

young are relatively well-developed when born

Ref­er­ences

Baker, R.H. 1983. Mam­mals of Michi­gan. Michi­gan State Uni­ver­sity Press.

Ban­field, A.W.F. 1981. The Mam­mals of Canada. Toronto Uni­ver­sity Press. Toronto, Buf­falo.

Carey, J., D. Judge. 2002. "Longevity Records: Life Spans of Mam­mals, Birds, Am­phib­ians, Rep­tiles, and Fish" (On-line). Max Planck In­sti­tute for De­mo­graphic Re­search. Ac­cessed May 18, 2007 at http://​www.​demogr.​mpg.​de/​.

Kurta, A. 1995. Mam­mals of the Great Lakes Re­gion. Ann Arbor: Uni­ver­sity of Michi­gan Press.

Wil­son, D., S. Ruff. 1999. The Smith­son­ian Book of North Amer­i­can Mam­mals. Wash­ing­ton, D.C.: The Smith­son­ian In­sti­tu­tion Press.