Hyracoideahyraxes

Hyraxes, also known as dassies or conies, are jack-rab­bit sized an­i­mals with short tails and pe­cu­liar, 3-toed hind feet with al­most hoof-like nails on two of the toes (the inner toe has a claw). The forefeet have 5 toes. The soles of their fore- and hind­feet, which are moist­ened by spe­cial sweat glands, are re­mark­ably soft and elas­tic, which works to in­crease their fric­tion against the sub­strate. They have spe­cial­ized mus­cles in the soles of the feet that help them to work al­most like a suc­tion cup. The feet of hyraxes are mesax­onic, mean­ing that the plane of sym­me­try of the foot goes through the third digit. In this they re­sem­ble peris­so­dactyls.

The skulls of hyraxes have a short ros­trum, promi­nent pos­tor­bital processes that form a pos­tor­bital bar, small bul­lae, a large jugal that con­tributes to the gle­noid fossa, and a broad plate-like an­gu­lar re­gion of the mandible. Hyraxes have a sin­gle in­cisor on each side of the upper jaw (two on the lower), fol­lowed by a di­astema -- an arrange­ment that looks very ro­dent-like. Like those of ro­dents, the in­cisors of hyraxes grow con­tin­u­ously and have enamel only on the an­te­rior sur­face, cre­at­ing a self-sharp­en­ing, chisel-like cut­ting edge. The mo­lars have strongly de­vel­oped lophs, as is com­mon in species with her­biv­o­rous diets. The den­tal for­mula is 1/2, 0/0, 4/4, 3/3 = 34.

Hyra­coids are usu­ally grouped with ele­phants and sire­ni­ans as "sub­un­gu­lates," and they all may have all de­scended from a com­mon stock

The diet of hyraxes con­sists mostly of leaves, bark and grasses, but they also eat some in­sects. Some species are ar­bo­real, while oth­ers live on rock out­crops. All are quick and agile climbers. They are ac­tive dur­ing day­light hours. Rock-dwelling species live in colonies; ar­bo­real species tend to be soli­tary. All make a va­ri­ety of whis­tles, chat­ters, and other sounds.

There is a sin­gle fam­ily of hyraxes, Pro­cavi­idae, and seven liv­ing species. Hyraxes are found in sub-Sa­ha­ran Africa and Mid­dle East.

Tech­ni­cal char­ac­ters

Lit­er­a­ture and ref­er­ences cited

Feld­hamer, G. A., L. C. Drick­amer, S. H. Vessey, and J. F. Mer­ritt. 1999. Mam­mal­ogy. Adap­ta­tion, Di­ver­sity, and Ecol­ogy. WCB Mc­Graw-Hill, Boston. xii+563pp.

Jones, C. 1984. Tubu­li­den­tates, pro­boscideans, and hyra­coideans. Pp. 523-535 in An­der­son, S. and J. K. Jones, Jr. (eds). Or­ders and Fam­i­lies of Re­cent Mam­mals of the World. John Wiley and Sons, N.Y. xii+686 pp.

Vaughan, T. A. 1986. Mam­mal­ogy. Third Edi­tion. Saun­ders Col­lege Pub­lish­ing, Fort Worth. vii+576 pp.

Vaughan, T. A., J. M. Ryan, N. J. Czaplewski. 2000. Mam­mal­ogy. Fourth Edi­tion. Saun­ders Col­lege Pub­lish­ing, Philadel­phia. vii+565pp.

Con­trib­u­tors

Phil Myers (au­thor), Mu­seum of Zo­ol­ogy, Uni­ver­sity of Michi­gan-Ann Arbor.

Glossary

bilateral symmetry

having body symmetry such that the animal can be divided in one plane into two mirror-image halves. Animals with bilateral symmetry have dorsal and ventral sides, as well as anterior and posterior ends. Synapomorphy of the Bilateria.

chemical

uses smells or other chemicals to communicate

endothermic

animals that use metabolically generated heat to regulate body temperature independently of ambient temperature. Endothermy is a synapomorphy of the Mammalia, although it may have arisen in a (now extinct) synapsid ancestor; the fossil record does not distinguish these possibilities. Convergent in birds.

motile

having the capacity to move from one place to another.

sexual

reproduction that includes combining the genetic contribution of two individuals, a male and a female

tactile

uses touch to communicate