Hesperia uncas

Ge­o­graphic Range

The Uncas skip­per, Hes­pe­ria uncas, ranges across Canada from Saskatchewan to Al­berta, and down into the United States through the Rocky Moun­tains and Great Plains. From the Great Plains its range ex­tends south to Texas and New Mex­ico.

There are two iso­lated pop­u­la­tions, one in south cen­tral Mex­ico and the other in Min­nesota. In Min­nesota, a small pop­u­la­tion only ex­ists in the sand dunes of Sher­burne County. This pop­u­la­tion is 500 kilo­me­ters east of its known lim­its in the Dako­tas. This pop­u­la­tion in Min­nesota is dif­fer­ent, phe­no­typ­i­cally, than that of the pop­u­la­tion in the Great Plains. Two re­cent sight­ings of H. uncas males in south­west­ern Min­nesota might in­di­cate a wider dis­tri­b­u­tion in the east­ern side of South Dakota, how­ever lack of habi­tat in this area would likely pre­vent the es­tab­lish­ment of a pop­u­la­tion. (Cof­fin and Pfan­n­muller, 1988)

Habi­tat

Uncas skip­pers can pri­mar­ily be found in both al­ka­line and short grass­lands, as well as grassy al­ka­line flats. Some west­ern pop­u­la­tions ap­pear com­monly clus­tered on this­tles in south­west­ern road ditches by the dozens. Uncas skip­pers in Cal­i­for­nia are lo­cated at higher el­e­va­tions in the White Moun­tains of Mono and Inyo. (Brock, 2003; Garth and Tilden, 1986; Glass­berg, 2001)

  • Range elevation
    989 to 8100 m
    3244.75 to 26574.80 ft
  • Average elevation
    5500 m
    18044.62 ft

Phys­i­cal De­scrip­tion

The Uncas skip­per is very sim­i­lar to other skip­pers that it flies with, but this but­ter­fly is small at 28 to 34 mm long, with a ro­bust body, an­gled forewings, and shorter, more round hind wings. Its wingspan av­er­ages 3.1 to 3.81 cm. On the un­der­side of its wings, the Uncas skip­per has ex­ten­sive hind­wings of white chevron with con­nected spots, with white veins and blotchy dark brown to black mark­ings. The white spots ori­en­tate in­wardly and out­wardly by dark green shades. The pop­u­la­tions lo­cated in Mono County, Cal­i­for­nia have re­duced hind­wing white vein­ing and often have dark brown to black mark­ings. Like all skip­pers, Uncas skip­pers have an­ten­nas that are rel­a­tively short which have clubbed ends and tips that curve back­wards.

Males and fe­males dif­fer greatly on the upper side of the wings. Males will typ­i­cally have a brown­ish or­ange darker col­oration. Small light col­ored spots can be seen near the forewing tips. Males also have a lin­ear black band of scent scales that are used in courtship. These scales are placed on the long axis of the forewings. Fe­males tend to be darker, with larger white spots on the forewings. Uncas fe­males tend to have larger, and more broad wings which is a char­ac­ter­is­tic found in skip­pers. Both sexes will be­come duller with age as the scales fall off, but the pat­tern will never dis­ap­pear.

The eggs ap­pear green­ish white and are hemi­spher­i­cal. The young lar­vae form is pale brown, and the head is a darker brown with cream spots and streaks in the front. ("Hes­pe­ria uncas", 2014; Cof­fin and Pfan­n­muller, 1988; Glass­berg, 2001; "Hes­pe­ria uncas", 2014; Opler, 1999)

  • Sexual Dimorphism
  • female larger
  • sexes colored or patterned differently
  • ornamentation
  • Range length
    28 to 34 mm
    1.10 to 1.34 in
  • Range wingspan
    3.3 to 3.81 cm
    1.30 to 1.50 in

De­vel­op­ment

Pupae emerge dur­ing the spring or sum­mer, de­pend­ing on the re­gion. In South Dakota and much of its range, H. uncas has two gen­er­a­tions, one in May and June, the other in July to Sep­tem­ber. In Min­nesota, it has only one gen­er­a­tion in June and July. After emerg­ing from pu­pa­tion, adults take flight and find mates. Eggs are laid, and hatch after about 10 days. In the first gen­er­a­tion, lar­vae go through sev­eral in­star stages, pu­pate, and then emerge as adults. In the sec­ond gen­er­a­tion (or for the only gen­er­a­tion in Min­nesota), lar­vae de­velop until the fourth or fifth in­star, and then go into hi­ber­na­tion for the win­ter. They emerge again in late spring or sum­mer, and pu­pate for 2 to 3 weeks be­fore emerg­ing as adults and be­gin­ning the cycle again. ("Hes­pe­ria uncas", 2014)

Re­pro­duc­tion

Based on sim­i­lar species in the genus, fe­male Uncas skip­pers prob­a­bly mate shortly after emer­gence and will rarely re­mate. The males will look for re­cep­tive fe­males by perch­ing on low veg­e­ta­tion or di­rectly on bare sand. Males use the scent scales on their wings to pro­duce pheromones for courtship. Many males will also col­lect on hills or ridges, which may draw the at­ten­tion of fe­males, though lit­tle is known for sure about this be­hav­ior. Re­cep­tive fe­males re­spond to males by de­scend­ing into the veg­e­ta­tion where mat­ing fol­lows. ("Hes­pe­ria uncas", 2014)

De­pend­ing on the area, Uncas skip­pers can have one or two gen­er­a­tions per year. In South Dakota and most of its range, there are two gen­er­a­tion per year, with adults pre­sent from May to June and July to Sep­tem­ber. In Min­nesota, there is one gen­er­a­tion per year from June to July. After mat­ing, the fe­male finds plants to oviposit on; all ob­served ovipo­si­tions of H. uncas have been made on hairy grama (Boutelous hir­sute), which is a tufted grass. The fe­male will lay only a sin­gle egg be­fore mov­ing on to the next tuft. (Glass­berg, 2001; "Hes­pe­ria uncas", 2014)

  • Breeding interval
    Females mate once in their lives, while males will mate as much as possible.
  • Breeding season
    Depending on the region, mating takes place from late spring to late summer.

Fe­males pro­vide pro­vi­sion­ing in their eggs, as well as lay the eggs on suit­able host plants for the lar­vae to feed upon hatch­ing. After the eggs are laid, there is no more in­ter­ac­tion or care by the par­ents. ("Hes­pe­ria uncas", 2014)

  • Parental Investment
  • pre-hatching/birth
    • provisioning
      • female

Lifes­pan/Longevity

Adults of Hes­pe­ria uncas likely live for about 3 weeks after emerg­ing from pu­pa­tion. ("Hes­pe­ria uncas", 2014)

  • Average lifespan
    Status: wild
    21 days

Be­hav­ior

The lar­vae will live in shel­ters that grow in size, that they them­selves built out of silk and plant ma­te­ri­als. The shel­ters are built big­ger and big­ger as the lar­vae grow too. Over­win­ter­ing lar­vae have also been found in shel­ters that have been build under the soil. Adult Uncas skip­pers are very rapid and strong fliers. They have a very fast wing beat that al­most ap­pears to be like a blur to human eyes. ("Hes­pe­ria uncas", 2014)

Com­mu­ni­ca­tion and Per­cep­tion

Male Uncas skip­pers have a lin­ear black band of scent scales on their wings that re­lease pheromones dur­ing courtship. Males also may com­mu­ni­cate with fe­males vi­su­ally, as the males col­lect in groups on high ridges be­fore fe­males, likely a vi­sual cue to fe­males. ("Hes­pe­ria uncas", 2014)

Food Habits

The lar­val food source for Uncas skip­per is blue grama (Bouteloua gra­cilis) grass and other grasses such as hairy grama (Bouteloua hir­sute). Uncas skip­per adults seek nec­tar and will feed from al­most any flow­ers avail­able, but golden aster (Het­erotheca vil­losa) and hairy puc­coon (Lithos­per­mum car­olin­iense) are the pre­ferred sources of nec­tar. (Cof­fin and Pfan­n­muller, 1988; "Hes­pe­ria uncas", 2014)

  • Plant Foods
  • leaves
  • nectar

Pre­da­tion

There is no in­for­ma­tion avail­able about the preda­tors of Hes­pe­ria uncas. As quick fliers, they are likely able to evade any ground preda­tors, as well as many air­borne preda­tors. ("Hes­pe­ria uncas", 2014)

Ecosys­tem Roles

Uncas skip­pers, like other but­ter­flies and moths feed on flow­ers nec­tar and there­fore help con­tribute to pol­li­na­tion of plants. ("Hes­pe­ria uncas", 2014)

  • Ecosystem Impact
  • pollinates

Eco­nomic Im­por­tance for Hu­mans: Pos­i­tive

There are no known pos­i­tive ef­fects of Hes­pe­ria uncas on hu­mans.

Eco­nomic Im­por­tance for Hu­mans: Neg­a­tive

There are no known ad­verse ef­fects of Hes­pe­ria uncas on hu­mans.

Con­ser­va­tion Sta­tus

Under the Min­nesota's List of En­dan­gered, Threat­ened, and Spe­cial Con­cern Species, Herspe­ria uncas is listed as en­dan­gered. This is due to the de­struc­tion of its habi­tat in that state, and only a small pop­u­la­tion in an iso­lated area re­mains there. Con­ser­va­tion ef­forts are un­der­way to pro­tect the pop­u­la­tion and its habi­tat in this area.

On a na­tional level, Hes­pe­ria uncas has no spe­cial con­ser­va­tion sta­tus. ("Hes­pe­ria uncas", 2014)

Con­trib­u­tors

Court­ney Chris­tensen (au­thor), Min­nesota State Uni­ver­sity Mankato, Robert Sorensen (ed­i­tor), Min­nesota State Uni­ver­sity, Mankato, An­gela Miner (ed­i­tor), An­i­mal Di­ver­sity Web Staff.

Glossary

Nearctic

living in the Nearctic biogeographic province, the northern part of the New World. This includes Greenland, the Canadian Arctic islands, and all of the North American as far south as the highlands of central Mexico.

World Map

bilateral symmetry

having body symmetry such that the animal can be divided in one plane into two mirror-image halves. Animals with bilateral symmetry have dorsal and ventral sides, as well as anterior and posterior ends. Synapomorphy of the Bilateria.

chemical

uses smells or other chemicals to communicate

desert or dunes

in deserts low (less than 30 cm per year) and unpredictable rainfall results in landscapes dominated by plants and animals adapted to aridity. Vegetation is typically sparse, though spectacular blooms may occur following rain. Deserts can be cold or warm and daily temperates typically fluctuate. In dune areas vegetation is also sparse and conditions are dry. This is because sand does not hold water well so little is available to plants. In dunes near seas and oceans this is compounded by the influence of salt in the air and soil. Salt limits the ability of plants to take up water through their roots.

diapause

a period of time when growth or development is suspended in insects and other invertebrates, it can usually only be ended the appropriate environmental stimulus.

ectothermic

animals which must use heat acquired from the environment and behavioral adaptations to regulate body temperature

fertilization

union of egg and spermatozoan

folivore

an animal that mainly eats leaves.

herbivore

An animal that eats mainly plants or parts of plants.

heterothermic

having a body temperature that fluctuates with that of the immediate environment; having no mechanism or a poorly developed mechanism for regulating internal body temperature.

hibernation

the state that some animals enter during winter in which normal physiological processes are significantly reduced, thus lowering the animal's energy requirements. The act or condition of passing winter in a torpid or resting state, typically involving the abandonment of homoiothermy in mammals.

internal fertilization

fertilization takes place within the female's body

metamorphosis

A large change in the shape or structure of an animal that happens as the animal grows. In insects, "incomplete metamorphosis" is when young animals are similar to adults and change gradually into the adult form, and "complete metamorphosis" is when there is a profound change between larval and adult forms. Butterflies have complete metamorphosis, grasshoppers have incomplete metamorphosis.

motile

having the capacity to move from one place to another.

native range

the area in which the animal is naturally found, the region in which it is endemic.

nectarivore

an animal that mainly eats nectar from flowers

oviparous

reproduction in which eggs are released by the female; development of offspring occurs outside the mother's body.

pheromones

chemicals released into air or water that are detected by and responded to by other animals of the same species

polygynous

having more than one female as a mate at one time

seasonal breeding

breeding is confined to a particular season

sedentary

remains in the same area

semelparous

offspring are all produced in a single group (litter, clutch, etc.), after which the parent usually dies. Semelparous organisms often only live through a single season/year (or other periodic change in conditions) but may live for many seasons. In both cases reproduction occurs as a single investment of energy in offspring, with no future chance for investment in reproduction.

sexual

reproduction that includes combining the genetic contribution of two individuals, a male and a female

sexual ornamentation

one of the sexes (usually males) has special physical structures used in courting the other sex or fighting the same sex. For example: antlers, elongated tails, special spurs.

suburban

living in residential areas on the outskirts of large cities or towns.

temperate

that region of the Earth between 23.5 degrees North and 60 degrees North (between the Tropic of Cancer and the Arctic Circle) and between 23.5 degrees South and 60 degrees South (between the Tropic of Capricorn and the Antarctic Circle).

terrestrial

Living on the ground.

tropical savanna and grassland

A terrestrial biome. Savannas are grasslands with scattered individual trees that do not form a closed canopy. Extensive savannas are found in parts of subtropical and tropical Africa and South America, and in Australia.

savanna

A grassland with scattered trees or scattered clumps of trees, a type of community intermediate between grassland and forest. See also Tropical savanna and grassland biome.

temperate grassland

A terrestrial biome found in temperate latitudes (>23.5° N or S latitude). Vegetation is made up mostly of grasses, the height and species diversity of which depend largely on the amount of moisture available. Fire and grazing are important in the long-term maintenance of grasslands.

visual

uses sight to communicate

Ref­er­ences

Min­nesota De­part­ment of Nat­ural Re­sources. 2014. "Hes­pe­ria uncas" (On-line). Species Pro­file: Min­nesota DNR. Ac­cessed April 16, 2014 at http://​www.​dnr.​state.​mn.​us/​rsg/​profile.​html?​action=elementDetail&​selectedElement=IILEP65010.

Brock, J. 2003. Kauf­man Field Guide to But­ter­flies of North Amer­ica. New York, United States of Amer­ica: Houghton Mif­flin Com­pany.

Cof­fin, B., L. Pfan­n­muller. 1988. Min­nesota's En­dan­gered Flora and Fauna. Canada: Uni­ver­sity of Min­nesota Press. Ac­cessed March 26, 2014 at http://​mnsu.​summon.​serialssolutions.​com/?​s.​q=uncas+skipper#!/​search?​ho=t&​q=uncas%20skipper&​l=en.

Garth, J., J. Tilden. 1986. Cal­i­for­nia But­ter­flies. United States of Amer­ica: Uni­ver­sity of Cal­i­for­nia Press.

Glass­berg, J. 2001. But­ter­flies Through Binoc­u­lars. New York, United States of Amer­ica: Ox­ford Uni­ver­sity Press, Inc..

Hol­land, J. 2003. Field Guide to But­ter­flies. Canada: Ster­ling Pub­lish­ing Co, Inc..

Opler, P. 1999. A Field Guide to West­ern But­ter­flies. New York: Houghton Mif­flin Com­pany. Ac­cessed April 21, 2004 at http://​books.​google.​com/​books?​id=ilL_​XX1rbNoC&​pg=RA2-PT360&​dq=Holland+field+guide+to+butterflies&​hl=en&​sa=X&​ei=MeJVU5y7HNGKyATKjYKIAw&​ved=0CEQQ6AEwAg#​v=onepage&​q=uncas%20skipper&​f=false.