Chordeiles minorcommon nighthawk

Ge­o­graphic Range

Com­mon Nighthawks (Chordeiles minor) breed through­out much of North Amer­ica and parts of Cen­tral Amer­ica. Their win­ter dis­tri­b­u­tion is less well known, but they are be­lieved to range through­out mid­dle South Amer­ica in the low­lands east of the Andes. (; Poulin, et al., 1996)

Habi­tat

Com­mon nighthawks breed­ing habi­tats in­clude coastal dunes and beaches, wood­land clear­ings, grass­lands, sa­van­nas, sage­brush plains, and open forests. They will also use habi­tat al­tered by human ac­tiv­ity in­clud­ing logged or burned areas of forests, farm fields, and cities.

Com­mon nighthawks choose nest sites on the ground in open areas with some cover from grasses, shrubs, logs, or boul­ders. They do not build nests. In­stead, eggs are laid on a va­ri­ety of sub­strates in­clud­ing sand, gravel, leaves, and bare rock. In areas of human habi­ta­tion, com­mon nighthawks often nest on flat, gravel roofs.

Lit­tle is known about the mi­gra­tion routes or win­ter habi­tat of com­mon nighthawks. They have been seen mi­grat­ing across wet­lands, farm­land, river val­leys, open wood­lands, and coastal dunes. They are pre­sumed to pre­fer open coun­try in their win­ter­ing sites and have been seen fly­ing over cities and towns. (; Poulin, et al., 1996; Stiles and Skutch, 1989)

  • Range elevation
    sea level (low) m
    (low) ft

Phys­i­cal De­scrip­tion

Com­mon nighthawks are medium-sized birds. They are 22 to 24 cm long and weigh 65 to 98 g. Like other mem­bers of the Caprimulgi­dae, they have large mouths and eyes, and are cryp­ti­cally col­ored. They have a notched tail and long, slen­der, pointed wings with white patches on the pri­maries. Males have a white tail band near the tip of the tail and a white throat patch. Fe­males do not have a tail band and are more buff-col­ored on the throat. Both sexes have bold bar­ring on the chest and belly, though light parts tend to be whiter on males and more buff-col­ored on fe­males.

Nine sub­species of com­mon nighthawks have been de­scribed. These are dif­fer­en­ti­ated by light and dark color vari­a­tions in the plumage. Com­mon nighthawks are often con­fused with two very sim­i­lar species of nighthawks: Lesser nighthawks (Chordeiles acu­tipen­nis) and An­til­lean nighthawks (Chordeiles gund­lachii). Lesser nighthawks are slightly smaller than Com­mon Nighthawks, with buffy un­der­tail-coverts in­stead of white, and with the white wing-patch of the pri­maries slightly closer to the wing tip. They also for­age closer to the ground than do com­mon nighthawks. An­til­lean nighthawks are vir­tu­ally in­dis­tin­guish­able from com­mon nighthawks in the field but by call, a nasal kil­likidick with the same tone as com­mon nighthawks’ peent. In the hand, An­til­lean nighthawks’ wing mea­sure­ments are slightly shorter than com­mon nighthawks’. (Ehrlich, 1988; ; Poulin, et al., 1996)

  • Sexual Dimorphism
  • sexes colored or patterned differently
  • Range mass
    65 to 98 g
    2.29 to 3.45 oz
  • Range length
    22 to 24 cm
    8.66 to 9.45 in
  • Average basal metabolic rate
    0.4421 W
    AnAge

Re­pro­duc­tion

There is lit­tle in­for­ma­tion avail­able about the mat­ing sys­tem of com­mon nighthawks. Males court fe­males by dis­play­ing on the ground and in flight. They begin by fly­ing 5 to 30 me­ters into the air and then div­ing steeply to­ward the ground, pulling up sharply about 2 me­ters above the ground. This dis­play is ac­com­pa­nied by a “boom­ing” noise made by the air rush­ing through the pri­mary feath­ers of the male. Males then land near the fe­male, spread­ing and shak­ing their tail from side to side, dis­play­ing their white throat patch, and mak­ing a croak­ing noise.

Com­mon nighthawk breed­ing phe­nol­ogy varies through­out their range, with more southerly pop­u­la­tions pro­duc­ing young as early as May and northerly pop­u­la­tions as late as Au­gust. Lit­tle is known about pair for­ma­tion or breed­ing ac­tiv­ity. Fe­male com­mon nighthawks have been shown to ar­rive first at their breed­ing grounds and se­lect the nest site. Banded in­di­vid­u­als have re­turned to the same nest sites over mul­ti­ple years. The age at first breed­ing is un­known for this species. Com­mon nighthawks are as­sumed to breed once per year.

Fe­males typ­i­cally lay 2 eggs, 1 to 2 days apart. The eggs are pale, splotched with gray, brown, and black. The fe­male in­cu­bates the eggs, leav­ing the nest in early evening to feed. In­cu­ba­tion pe­ri­ods vary through­out the breed­ing range be­tween 18-20 days. After the young hatch, the fe­male con­tin­ues to leave the nest site to for­age in the evening. She feeds re­gur­gi­tated in­sects to the young be­fore sun­rise in the morn­ing and after sun­set in the evening. Nestlings are semi­pre­co­cial and able to move in re­sponse to the fe­male’s calls within a day of hatch­ing. The young can move to shade or sun to reg­u­late their body tem­per­a­ture. After 16 days, young can hop. At 18 days they make their first flight, and can fly well at about 25-30 days old. By the time they are 30 days old, chicks have left the nest for good. Full de­vel­op­ment is achieved at 45-50 days, after which young may join mi­grat­ing flocks. In south­ern parts of the breed­ing range, pairs may have a sec­ond brood. In this case, the male takes over feed­ing the young of the first clutch while the fe­male in­cu­bates the sec­ond clutch. He will also feed the fe­male. (; Poulin, et al., 1996)

  • Breeding season
    spring and summer
  • Range eggs per season
    1 to 2
  • Average eggs per season
    2
  • Average eggs per season
    2
    AnAge
  • Average time to hatching
    18-20 days
  • Average time to hatching
    19 days
    AnAge
  • Average fledging age
    18 days
  • Average time to independence
    30 days

The fe­male of a breed­ing pair in­cu­bates the eggs and broods the young chicks. Though the young are able to move them­selves only one day after hatch­ing, the fe­male may move them around to take ad­van­tage of nearby shade. The par­ents feed the chicks re­gur­gi­tated in­sects until they are able to feed them­selves at age 25 days or so. The male de­fends the nest site by wing-beat­ing and hiss­ing at in­trud­ers. The fe­male may also de­fend the nest site by feign­ing in­jury or hiss­ing at an in­truder. (; Poulin, et al., 1996)

  • Parental Investment
  • precocial
  • pre-fertilization
    • provisioning
    • protecting
      • male
      • female
  • pre-hatching/birth
    • protecting
      • male
      • female
  • pre-weaning/fledging
    • provisioning
      • male
      • female
    • protecting
      • male
      • female

Lifes­pan/Longevity

Com­mon nighthawks are ex­pected to live at least 4 to 5 years. The old­est known wild com­mon nighthawk was 9 years old. (; Poulin, et al., 1996)

  • Range lifespan
    Status: wild
    9 (high) years
  • Typical lifespan
    Status: wild
    4 to 5 years
  • Average lifespan
    Status: wild
    4 to 5 years

Be­hav­ior

Com­mon nighthawks are some of the last mi­gra­tory birds to ar­rive at spring breed­ing grounds and some of the first to leave in fall. This is at­trib­uted to the fact that warm tem­per­a­tures are req­ui­site for fly­ing in­sect ac­tiv­ity in early evening and morn­ing. Also, while some caprimulgids go into tor­por to con­serve en­ergy on cold nights, com­mon nighthawks are not known to do so, thus mak­ing them more sen­si­tive to colder tem­per­a­tures.

Com­mon nighthawks mi­grate some 4,000 to 11,000 km (2,500 to 6,800 miles) – one of the longest mi­gra­tions of any bird in the Amer­i­cas. They mi­grate in flocks, es­pe­cially in the fall, most pass­ing through Mid­dle Amer­ica, but some cross­ing the Gulf of Mex­ico and Caribbean to Florida. They may be seen rid­ing ther­mals at this time, much like mi­grat­ing hawks.

Com­mon nighthawks are highly ter­ri­to­r­ial, soli­tary nesters. Males will de­fend their ter­ri­tory by div­ing at in­trud­ers. The ter­ri­tory size varies be­tween habi­tat types, but ap­pears to range be­tween 41,000 and 280,000 square me­ters.

Com­mon nighthawks are cre­pus­cu­lar. When not at the nest, Com­mon nighthawks will roost, sleep, or sun­bathe on fence posts, tree branches, the ground, or flat rooftops.

Bats and lesser nighthawks have been shown to dis­place com­mon nighthawks from feed­ing sites by chas­ing them away. Lesser nighthawk ag­gres­sion may ex­clude com­mon nighthawks from mu­tu­ally suit­able habi­tats. (El­ph­ick, 1995; ; Poulin, et al., 1996)

  • Range territory size
    41000 to 28000 m^2

Com­mu­ni­ca­tion and Per­cep­tion

Com­mon nighthawks use calls and dis­plays to com­mu­ni­cate with one an­other. The vo­cal­iza­tions of com­mon nighthawks are very sim­ple, and have few vari­a­tions. They also use non-vo­cal sounds, such as the boom­ing sound made by the pri­mary feath­ers of males dur­ing a courtship dis­play to com­mu­ni­cate. An ex­am­ple of the phys­i­cal dis­plays used by com­mon nighthawks is the div­ing dis­play given by males to prospec­tive mates. (; Poulin, et al., 1996)

Food Habits

Com­mon nighthawks are cre­pus­cu­lar. They are most ac­tive at dawn and dusk, and rarely feed at night. They have been re­ported to oc­ca­sion­ally feed dur­ing the day in low light con­di­tions (stormy weather or fog, for ex­am­ple) They use their large mouths to “hawk” in­sects in the air. Their large eyes help them find and dis­tin­guish among prey items in the dark. Like owls, com­mon nighthawks have a tape­tum (a mir­ror-like struc­ture at the back of each eye that re­flects light to the retina) that in­creases their abil­ity to see in the dark. They fly with er­ratic, bat-like move­ments, tak­ing as many as 50 dif­fer­ent in­sect prey species. Stud­ies in­di­cate that the ma­jor­ity of the diet is made up of queen ants (Hy­menoptera), bee­tles (Coleoptera), and true bugs (Ho­moptera). It also in­cludes moths (Lep­i­doptera), mayflies (Ephemeroptera), cad­dis­flies (Tri­choptera), flies (Diptera), wasps (Hy­menoptera), crick­ets and grasshop­pers (Or­thoptera) and other in­sects. In the urban parts of their range, com­mon nighthawks are often seen fly­ing around street­lights or bright yard lights, catch­ing in­sects that are at­tracted to the light.

Com­mon Nighthawks drink while in flight by skim­ming the sur­face of lakes, streams, or water troughs with their bills. (Brigham and Bar­clay, 1995; Brigham, 1990; Nicol and Arnott, 1974; ; Poulin, et al., 1996; Ter­res, 1980)

  • Animal Foods
  • insects

Pre­da­tion

Crows and ravens, owls, Amer­i­can kestrels, skunks, snakes, house­cats and canids such as coy­otes, foxes, and do­mes­tic dogs are preda­tors of com­mon nighthawks.

Fe­males and young rely on their cryp­tic brown col­oration to avoid de­tec­tion of the nest site by preda­tors. Males are not known to guard the nest but will de­fend it by div­ing over it and boom­ing with their wings or beat­ing the wings and hiss­ing. Fe­males may feign in­jury to dis­tract preda­tors from the nest. Chicks also spread their wings and hiss at in­trud­ers when threat­ened. (; Poulin, et al., 1996)

  • Anti-predator Adaptations
  • cryptic

Ecosys­tem Roles

Com­mon nighthawks help to con­trol pop­u­la­tions of the in­sects that they prey on. They also com­pete with bats and lesser nighthawks Chordeiles acu­tipen­nis for food. (; Poulin, et al., 1996)

Eco­nomic Im­por­tance for Hu­mans: Pos­i­tive

Since com­mon nighthawks are in­sect eaters that fre­quent farm fields and cities, it is likely that they help con­trol pest in­sect species.

  • Positive Impacts
  • controls pest population

Eco­nomic Im­por­tance for Hu­mans: Neg­a­tive

Com­mon nighthawks have no known neg­a­tive im­pact on hu­mans.

Con­ser­va­tion Sta­tus

Pop­u­la­tions of com­mon nighthawks are de­clin­ing. This de­crease may be at­trib­uted to a va­ri­ety of human ac­tiv­i­ties. In­dis­crim­i­nate pes­ti­cide use in cities and farm­lands af­fects pop­u­la­tions lo­cally. In urban areas, re­place­ment of gravel roofs with rub­ber roofs has re­duced nest­ing sites for these pop­u­la­tions. In­creased pre­da­tion is also a fac­tor in gen­eral pop­u­la­tion de­cline. Urban nesters are es­pe­cially vul­ner­a­ble to pre­da­tion by house­cats. Com­mon nighthawks are also killed by ve­hi­cles when roost­ing or feed­ing along road­ways. (Poulin, et al., 1996)

Other Com­ments

Per­sonal anec­dote:

Com­mon Nighthawks are known for their loud, nasal “peent” calls made by both sexes, as well as the males’ amaz­ing, boom­ing courtship dives. My hus­band and I were equally freaked-out and cap­ti­vated by their mys­te­ri­ous sounds one sum­mer night in east­ern Ari­zona. We were dri­ving home from Col­orado, and the sun had long-since gone down. Eye­lids heavy, we pulled off onto na­tional for­est land just east of Show Low to camp for the night. We laid our tarp and sleep­ing bags on rough vol­canic grav­els in a sparse ju­niper wood­land. There was no moon. Just as we had fallen asleep, we were awaken by a loud “peent­ing” noise, fol­lowed by a boom­ing, zip­ping “woosh.” It passed right over our heads. A few sec­onds later, the peent came again from a com­pletely dif­fer­ent cor­ner of the sky, fol­lowed by an­other close woosh above our heads. Our first thought was that bored teenagers from Show Low had some­how found our camp­site and were mess­ing with our minds. What WAS that sound?!? Strangely muf­fled gun­shots? UFO’s land­ing? A huge bug-zap­per? It just didn’t sound nat­ural. How could some­thing be in one spot, then abruptly be 200 me­ters away, mak­ing such a mech­a­nized sound? Then, the sound ceased, leav­ing us to won­der its ori­gins. We drifted off to sleep, only to be awoken a while be­fore dawn.

Peent! Woosh! Si­lence. Peent… again from a spot im­pos­si­bly dis­tant from the first call…and woosh above our heads. Dis­be­lief gave way to rea­son as we hun­kered in our bags, com­mis­er­at­ing: “It’s got to be a bird.” “Maybe some kind of night­jar?” Sure enough, as the sun grad­u­ally light­ened the east­ern sky, we began to make out an avian shape. It would flap up on slen­der, pointed wings, hover, give out a loud “peent”, and dive steeply. The me­chan­i­cal woosh and zip­ping noise came with the dive and ceased with the bird’s abrupt re­turn sky­ward. It would peent again a cou­ple hun­dred me­ters from where it began its last dive, then plunge down­ward. Boom-woosh! As the sky light­ened, we saw the flash of white wing-patches on a sec­ond bird – per­haps a fe­male? Then they moved off and were gone. We con­sulted our bird books later and de­cided they must have been com­mon nighthawks. Could they have been court­ing? Were we dis­turb­ing a nest site? We’ll never know. Though no records of night­time div­ing dis­plays exist for this species, we def­i­nitely wit­nessed them late into the night and well be­fore dawn. An un­for­get­table ex­pe­ri­ence!

Con­trib­u­tors

Kari Kirschbaum (ed­i­tor), An­i­mal Di­ver­sity Web.

Robin Kropp (au­thor), Uni­ver­sity of Ari­zona, Jorge Schon­dube (ed­i­tor), Uni­ver­sity of Ari­zona.

Glossary

Nearctic

living in the Nearctic biogeographic province, the northern part of the New World. This includes Greenland, the Canadian Arctic islands, and all of the North American as far south as the highlands of central Mexico.

World Map

Neotropical

living in the southern part of the New World. In other words, Central and South America.

World Map

acoustic

uses sound to communicate

agricultural

living in landscapes dominated by human agriculture.

bilateral symmetry

having body symmetry such that the animal can be divided in one plane into two mirror-image halves. Animals with bilateral symmetry have dorsal and ventral sides, as well as anterior and posterior ends. Synapomorphy of the Bilateria.

carnivore

an animal that mainly eats meat

chemical

uses smells or other chemicals to communicate

crepuscular

active at dawn and dusk

cryptic

having markings, coloration, shapes, or other features that cause an animal to be camouflaged in its natural environment; being difficult to see or otherwise detect.

desert or dunes

in deserts low (less than 30 cm per year) and unpredictable rainfall results in landscapes dominated by plants and animals adapted to aridity. Vegetation is typically sparse, though spectacular blooms may occur following rain. Deserts can be cold or warm and daily temperates typically fluctuate. In dune areas vegetation is also sparse and conditions are dry. This is because sand does not hold water well so little is available to plants. In dunes near seas and oceans this is compounded by the influence of salt in the air and soil. Salt limits the ability of plants to take up water through their roots.

endothermic

animals that use metabolically generated heat to regulate body temperature independently of ambient temperature. Endothermy is a synapomorphy of the Mammalia, although it may have arisen in a (now extinct) synapsid ancestor; the fossil record does not distinguish these possibilities. Convergent in birds.

forest

forest biomes are dominated by trees, otherwise forest biomes can vary widely in amount of precipitation and seasonality.

insectivore

An animal that eats mainly insects or spiders.

iteroparous

offspring are produced in more than one group (litters, clutches, etc.) and across multiple seasons (or other periods hospitable to reproduction). Iteroparous animals must, by definition, survive over multiple seasons (or periodic condition changes).

migratory

makes seasonal movements between breeding and wintering grounds

monogamous

Having one mate at a time.

motile

having the capacity to move from one place to another.

native range

the area in which the animal is naturally found, the region in which it is endemic.

oviparous

reproduction in which eggs are released by the female; development of offspring occurs outside the mother's body.

polymorphic

"many forms." A species is polymorphic if its individuals can be divided into two or more easily recognized groups, based on structure, color, or other similar characteristics. The term only applies when the distinct groups can be found in the same area; graded or clinal variation throughout the range of a species (e.g. a north-to-south decrease in size) is not polymorphism. Polymorphic characteristics may be inherited because the differences have a genetic basis, or they may be the result of environmental influences. We do not consider sexual differences (i.e. sexual dimorphism), seasonal changes (e.g. change in fur color), or age-related changes to be polymorphic. Polymorphism in a local population can be an adaptation to prevent density-dependent predation, where predators preferentially prey on the most common morph.

rainforest

rainforests, both temperate and tropical, are dominated by trees often forming a closed canopy with little light reaching the ground. Epiphytes and climbing plants are also abundant. Precipitation is typically not limiting, but may be somewhat seasonal.

scrub forest

scrub forests develop in areas that experience dry seasons.

seasonal breeding

breeding is confined to a particular season

sexual

reproduction that includes combining the genetic contribution of two individuals, a male and a female

solitary

lives alone

suburban

living in residential areas on the outskirts of large cities or towns.

tactile

uses touch to communicate

temperate

that region of the Earth between 23.5 degrees North and 60 degrees North (between the Tropic of Cancer and the Arctic Circle) and between 23.5 degrees South and 60 degrees South (between the Tropic of Capricorn and the Antarctic Circle).

terrestrial

Living on the ground.

territorial

defends an area within the home range, occupied by a single animals or group of animals of the same species and held through overt defense, display, or advertisement

tropical

the region of the earth that surrounds the equator, from 23.5 degrees north to 23.5 degrees south.

tropical savanna and grassland

A terrestrial biome. Savannas are grasslands with scattered individual trees that do not form a closed canopy. Extensive savannas are found in parts of subtropical and tropical Africa and South America, and in Australia.

savanna

A grassland with scattered trees or scattered clumps of trees, a type of community intermediate between grassland and forest. See also Tropical savanna and grassland biome.

temperate grassland

A terrestrial biome found in temperate latitudes (>23.5° N or S latitude). Vegetation is made up mostly of grasses, the height and species diversity of which depend largely on the amount of moisture available. Fire and grazing are important in the long-term maintenance of grasslands.

urban

living in cities and large towns, landscapes dominated by human structures and activity.

visual

uses sight to communicate

young precocial

young are relatively well-developed when born

Ref­er­ences

Brigham, R. 1990. Prey se­lec­tion by big brown bats (Eptesi­cus fus­cus) and Com­mon Nighthawks (Chordeiles minor). Amer­i­can Mid­land Nat­u­ral­ist, 124: 73-80.

Brigham, R., R. Bar­clay. 1995. Prey se­lec­tion by Com­mon Nighthawks: does vi­sion im­pose a con­straint?. Eco­science, 2(3): 276-279.

Ehrlich, P. 1988. A Birder’s Hand­book: a field guide to the nat­ural his­tory of North Amer­i­can birds. New York: Simon and Schus­ter.

El­ph­ick, J. 1995. Atlas of Bird Mi­gra­tion. Lon­don: Harper-Collins Pub­lish­ers, Ltd.

Nicol, J., H. Arnott. 1974. Tapeta lu­cidum in the eyes of goat­suck­ers (Caprimulgi­dae). Pro­ceed­ings of the Royal Acad­emy of Lon­don, 187: 349-352.

Poulin, R., S. Grindal, R. Brigham. 1996. Com­mon Nighthawk (Chordeiles minor). A Poole, F Gill, eds. The Birds of North Amer­ica, Vol. 213. Philadel­phia, PA and Wash­ing­ton DC: The Acad­emy of Nat­ural Sci­en­tists and The Amer­i­can Or­nithol­o­gists Union.

Stiles, F., A. Skutch. 1989. A Guide to the Birds of Costa Rica. Ithaca: Cor­nell Uni­ver­sity Press.

Ter­res, J. 1980. The Audubon so­ci­ety en­cy­clo­pe­dia of North Amer­i­can birds. New York: Al­fred A. Knopf.