Aythya fuligulatufted duck

Ge­o­graphic Range

Tufted ducks are mi­gra­tory birds with a broad ge­o­graphic range that ex­tends from Eura­sia and Africa to the North Amer­i­can coasts. They were his­tor­i­cally na­tive only to the Palearc­tic Re­gion. How­ever, over the past cen­tury, their range has ex­panded due to an in­creased avail­abil­ity of open water due to man-made changes in its habi­tat. Man-made lakes have pro­vided these birds with ideal feed­ing habi­tats. (Bent, 1951; BirdLife In­ter­na­tional, 2011)

Dur­ing breed­ing months (May to early Au­gust), tufted ducks in­habit most Palearc­tic re­gions such as the Faroe Is­lands, the British Isles, Nor­way, nearly all of Eu­rope, and Asia to east­ern Siberia. The limit of their range ex­tends from about 70 de­grees North and South lat­i­tudes to about 50 de­grees north. Dur­ing the win­ter months (No­vem­ber to April), tufted ducks can be found in south­ern Eu­rope, north­ern Africa, south­ern Asia, and some areas through­out North Amer­ica. They can also be found oc­ca­sion­ally through­out areas of Madeira, Bor­neo, Liberia, the Sey­chelle, Pelew, Mar­i­anne and Philip­pine Is­lands. (Bent, 1951; BirdLife In­ter­na­tional, 2011)

Habi­tat

The habi­tat of tufted ducks varies sea­son­ally due to its mi­gra­tory be­hav­ior. Through­out the breed­ing sea­son, they are most often found in shal­low lakes. They pre­fer shal­low water rang­ing from 3 to 5 m deep, with tall thick wet­land veg­e­ta­tion, such as reeds, for perch­ing and preen­ing. Veg­e­ta­tion is also an im­por­tant fac­tor in pro­tec­tion from the wind. Dur­ing the breed­ing sea­son tufted ducks typ­i­cally avoid lakes that are deeper than 15 m. Dur­ing win­ter months, they can be found gen­er­ally in larger bod­ies of open water such as marshes, lakes, es­tu­ar­ies, and man-made ponds. Dur­ing pe­ri­ods of mi­gra­tion, they can also be found in and along rivers. (Bent, 1951; BirdLife In­ter­na­tional, 2009)

  • Aquatic Biomes
  • lakes and ponds
  • rivers and streams
  • coastal
  • Range depth
    2 to 15 m
    6.56 to 49.21 ft
  • Average depth
    7 m
    22.97 ft

Phys­i­cal De­scrip­tion

Tufted ducks are small to medium-sized div­ing ducks. Sex­ual di­mor­phism is clearly ap­par­ent among males and fe­males. Males are typ­i­cally larger than fe­males, with an av­er­age body length of 42 to 48 cm and a mass of 753.0 to 1026.2 g. Fe­males gen­er­ally range from 39 to 44 cm in length and weigh in at 629.8 to 906.8 g. Sea­son­ally, body mass will fluc­tu­ate be­tween sexes. Both male and fe­male tufted ducks have an av­er­age wingspan of 70 cm. (Cleeves, 2002; Dick, 2002; Robin­son, 2005)

The defin­ing char­ac­ter­is­tic of this species that dis­tin­guishes them from the other mem­bers of Anati­dae is the dis­tinct tuft lo­cated on the back of the head. The tuft is black and more promi­nent on males, and gen­er­ally brown and less no­tice­able on fe­males. Adult males are solid black with white un­der­bel­lies and flanks. They have bright yel­low eyes and a dis­tinct tuft that is some­times mat­ted down from div­ing. Adult fe­males are brown with dark yel­low flanks and a white un­der­belly and a less pro­nounced tuft, or some­times no tuft at all. Fe­males also have bright yel­low eyes. The wing un­der­sides of both adult fe­males and males is white. (Cleeves, 2002; Dick, 2002; Robin­son, 2005)

Ju­ve­nile tufted ducks are sim­i­lar in ap­pear­ance to ma­ture fe­male adults, how­ever, ju­ve­nile color is less vi­brant and the tuft is less pro­nounced. Males in non-breed­ing plumage also re­sem­ble fe­males with a brown tint and a less promi­nent tuft or no tuft at all. (Cleeves, 2002; Dick, 2002; Robin­son, 2005)

  • Sexual Dimorphism
  • male larger
  • sexes colored or patterned differently
  • male more colorful
  • Range mass
    753.0 (male) 629.8 (female) to 1026.2 (male) 906.8 (female) g
    to oz
  • Average mass
    889.6 (male) 768.3 (female) g
    oz
  • Range length
    406.4 to 457.2 mm
    16.00 to 18.00 in
  • Average length
    431.8 mm
    17.00 in
  • Range wingspan
    201.5 (male) 193.6 (female) to 212.1 (male) 206.6 (female) mm
    to in

Re­pro­duc­tion

Tufted ducks are monog­a­mous and breeds once a year. Adults form pair bonds dur­ing spring mi­gra­tion and mates re­main to­gether until late June to early July. Re­pro­duc­tive be­hav­ior and man­ner­isms are sim­i­lar to other div­ing ducks. Ex­am­ples of "show­ing off" be­hav­ior in­clude dip­ping, also known as drink­ing, which is when the males and fe­males swim quickly along­side one an­other and dip their bills in and out of the water. An­other ex­am­ple is the neck stretch, which is when the male swims rapidly past the fe­male and reaches its neck to its en­tire length, hold­ing it there for a few sec­onds. Male-only dis­plays are few but con­sist of nod swim­ming (a fast swim that in­cludes nod­ding the head back in forth with full ex­ten­sion), a head throw (sim­i­lar to neck stretch), and preen­ing be­hind the wing. (Bent, 1951; Cleeves, 2002; Robin­son, 2005)

The breed­ing sea­son for tufted ducks oc­curs from May through early Au­gust, with peak ac­tiv­ity from mid-May to mid-July. The breed­ing sea­son be­gins with pair bond­ings which are formed dur­ing spring mi­gra­tion and con­tinue until late June or early July. (Bent, 1951; Cleeves, 2002; Robin­son, 2005)

For nest site se­lec­tion, pairs browse along the open wa­ters of wet­lands. Fe­males swim into flooded areas look­ing for a suit­able nest­ing site while males stay alert for preda­tors and other threats. Ideal nest­ing areas are near water, gen­er­ally amid dense veg­e­ta­tion. The fe­male is the con­struc­tor of the nest which re­quires al­most a week to com­plete. Grasses and feath­ers are used as sup­port for the nest. By the third day of nest build­ing, a bowl shape has been formed; by day six, the nest is strong with a de­fined shape. (Bent, 1951; Cleeves, 2002; Robin­son, 2005)

Fe­male tufted ducks lay be­tween 6 and 14 eggs a sea­son, but on av­er­age be­tween 8 and 10. Eggs are smooth and ovu­lar in shape and range in color from olive-brown to olive-gray. In­cu­ba­tion starts after the clutch is com­pleted and be­gins on the first night the fe­male re­mains in the nest overnight. In­cu­ba­tion usu­ally lasts 26 to 27 days. Chicks hatch usu­ally weigh­ing 28 to 31 g, cov­ered in downy feath­ers, and can gen­er­ally begin fol­low­ing par­ents and feed­ing them­selves soon after hatch­ing. Fledg­ing oc­curs 49 to 56 days after hatch­ing and in­de­pen­dence oc­curs 21 to 56 days after fledg­ing. Re­pro­duc­tive ma­tu­rity is quickly reached by both sexes, and breed­ing can occur dur­ing the fol­low­ing breed­ing sea­son. (Bent, 1951; Cleeves, 2002; Robin­son, 2005)

  • Breeding interval
    Tufted ducks breed once yearly.
  • Breeding season
    The breeding season occurs during late winter or spring months
  • Range eggs per season
    8 to 11
  • Average eggs per season
    9 to 10
  • Range time to hatching
    25 to 29 days
  • Average time to hatching
    26 to 27 days
  • Range fledging age
    49 to 56 days
  • Range time to independence
    21 to 56 days
  • Average time to independence
    43 days
  • Average age at sexual or reproductive maturity (female)
    1 years
  • Average age at sexual or reproductive maturity (male)
    1 years

Parental care for duck­lings is ex­hib­ited only by fe­males. Males do con­tribute in­di­rectly by tak­ing care of nu­tri­tional needs of the fe­male dur­ing the lay­ing process. The young emerge from the egg with­out any help from the mother, after which she dis­poses of the egg by in­gest­ing it, re­mov­ing it from nest area, or crush­ing the shells. Young tufted ducks do not de­pend on their moth­ers for food after they learn to dive. This typ­i­cally oc­curs within the first 48 hours of hatch­ing, how­ever, dur­ing this first week young duck­lings rely mostly on sur­face foods. (Bent, 1951)

  • Parental Investment
  • precocial
  • female parental care
  • pre-fertilization
    • provisioning
    • protecting
      • male
      • female
  • pre-hatching/birth
    • provisioning
      • female
    • protecting
      • female
  • pre-weaning/fledging
    • protecting
      • female
  • pre-independence
    • protecting
      • female

Lifes­pan/Longevity

Lit­tle is known about the lifes­pan and longevity of tufted ducks.

Be­hav­ior

Tufted ducks are mi­gra­tory birds that travel sea­son­ally be­tween the breed­ing and over­win­ter­ing grounds. Like most ducks, they are highly aquatic and rarely stray far from a body of water. They are di­ur­nal for most of their life but may mi­grate noc­tur­nally. (Cleeves, 2002; Hill, 1983)

Dur­ing the breed­ing sea­son, male tufted duckss es­tab­lish ter­ri­to­ries sur­round­ing their nest and will de­fend it against neigh­bor­ing males and preda­tors. Dur­ing mi­gra­tory pe­ri­ods and the non-breed­ing sea­son, they are so­cial. Groups typ­i­cally gather in shal­low lakes, ponds, and slow mov­ing rivers. Males are gen­er­ally silent ex­cept dur­ing mat­ing. Fe­males make a "karr" sound dur­ing flight. (Cleeves, 2002; Hill, 1983)

This species can be dis­tin­guished from other div­ing ducks by their unique div­ing be­hav­ior. They launch them­selves slightly out of the water to help sub­merge their bod­ies un­der­wa­ter while for­ag­ing in shal­low water. Sub­mer­gence time for ju­ve­niles is less than that of adults. Duck­lings and ju­ve­niles skim the sur­face of wa­ters for emerg­ing in­sects and dive in very shal­low water for New Zealand mud snails (Potam­mopy­r­gus jenk­insi) and Zebra mus­sels (Dreis­sena poly­mor­pha). Sub­mer­gence time grad­u­ally in­creases as in­di­vid­u­als ma­ture and in­crease in size. (Cleeves, 2002; Hill, 1983)

Home Range

The range of tufted ducks for both breed­ing and res­i­dency is es­ti­mated at 20,400,000 km². Spe­cific ter­ri­tory sizes for male tufted ducks is cur­rently un­known. (BirdLife In­ter­na­tional, 2009)

Com­mu­ni­ca­tion and Per­cep­tion

The use of vi­sual and acoustic per­cep­tions is key for tufted ducks be­cause they are needed to com­mu­ni­cate for mat­ing rit­u­als and warn­ing calls. Like most birds, they per­ceive their en­vi­ron­ment through vi­sual, au­di­tory, tac­tile and chem­i­cal stim­uli. (Bent, 1951)

Both fe­males and males make sim­i­lar calls, korr,korr,korr, or ka-ka-ka, karr, with the fe­male nor­mally pre­sent­ing the call louder. Calls are nor­mally emit­ted when be­gin­ning to fly, quar­rel­ing, or when star­tled. (Bent, 1951)

Dur­ing mate se­lec­tion, two or three male ducks will flaunt their at­trib­utes in the pres­ence of a fe­male. To at­tract mates, males will swim in cir­cles around a fe­male, pass­ing her rapidly. The male will ex­tend his neck to full length and raises his bill but will not look in the fe­male's di­rec­tion. Like other div­ing ducks, male tufted ducks dip their bills fre­quently and sound calls dur­ing courtship. Fe­males have also been spot­ted flaunt­ing their at­trib­utes to­wards males be­fore mat­ing. (Bent, 1951)

Food Habits

Tufted ducks are om­ni­vores whose main source of food are mol­luscs (Mol­lusca). The most preyed upon mol­luscs are zebra mus­sels (Dreis­sena poly­mor­pha). Zebra mus­sels are gen­er­ally very abun­dant in slow mov­ing rivers, canals, docks, reser­voirs, and large areas of fresh­wa­ter. In ad­di­tion, tufted ducks con­sume a va­ri­ety of plant ma­te­ri­als, mostly leaves, stems, and roots. They also oc­ca­sion­ally feed on seeds. They gen­er­ally for­age and feed with other mem­bers of their species. They typ­i­cally dive to­gether, or one after the other very quickly, and stay sub­merged from a few sec­onds to one minute. (Bent, 1951; Olney, 1963)

There are al­ways quan­ti­ties of sand, fine shells, and small stones found within tufted ducks' stom­achs. This species usu­ally con­sumes food un­der­wa­ter; how­ever, when a larger ver­te­brate or plant is being con­sumed they bring that food to the sur­face where they crush the food with their bills be­fore in­gested. (Bent, 1951; Olney, 1963)

  • Animal Foods
  • amphibians
  • fish
  • insects
  • mollusks
  • Plant Foods
  • leaves
  • roots and tubers
  • seeds, grains, and nuts
  • bryophytes

Pre­da­tion

The main preda­tors of tufted ducks are hu­mans. They are pro­tected by hunt­ing laws dur­ing the breed­ing sea­son (May through early Au­gust); how­ever, they may be hunted dur­ing the rest of the year. They can also be hunted by large birds of prey such as hawks, as well as ter­res­trial preda­tors in­clud­ing foxes, rac­coons, and com­mon snap­ping tur­tles. In ad­di­tion, many an­i­mals prey on their eggs, in­clud­ing do­mes­tic dogs, crows, and skunks. Like many birds, fe­male tufted ducks ex­hibit cryp­tic col­oration to cam­ou­flage them­selves while in­cu­bat­ing the clutch. (Bent, 1951; Cleeves, 2002)

  • Anti-predator Adaptations
  • cryptic
  • Known Predators

Ecosys­tem Roles

Adult tufted ducks, eggs, and young are all com­mon sources of food for many preda­tors. In ad­di­tion, they serve as hosts to a species of avian nasal par­a­site. This par­a­site is found in mol­lusks as an in­ter­me­di­ate host, and then are con­sumed by tufted ducks. They are preda­tors of many aquatic in­ver­te­brates, and gen­er­ally have a sig­nif­i­cant im­pact on these pop­u­la­tions. (Bent, 1951; BirdLife In­ter­na­tional, 2011; Cleeves, 2002; Olney, 1963; Rudol­fova, et al., 2002)

Com­men­sal/Par­a­sitic Species
  • avian nasal par­a­sites (Tri­chol­bil­harazia re­genti)

Eco­nomic Im­por­tance for Hu­mans: Pos­i­tive

Tufted ducks are eco­nom­i­cally im­por­tant for the hunt­ing and game in­dus­try. The only time when the species can­not be hunted is dur­ing breed­ing sea­son. They pro­vide mo­ti­va­tion to con­serve wet­land habi­tats, which are of great eco­nomic im­por­tance to hu­mans. (Miller and Spool­man, 2008)

  • Positive Impacts
  • food

Eco­nomic Im­por­tance for Hu­mans: Neg­a­tive

There are no known neg­a­tive im­pacts of tufted ducks on hu­mans.

Con­ser­va­tion Sta­tus

Tufted ducks have a very large ge­o­graphic range and their pop­u­la­tions are not de­clin­ing or fluc­tu­at­ing. The con­ser­va­tion sta­tus of tufted ducks on the IUCN Red list is of least con­cern. The largest threat to tufted ducks is habi­tat loss through human in­duced al­ter­ation. This in­cludes de­struc­tion of wet­lands for human de­vel­op­ment, waste pol­lu­tion, and oil spills. (BirdLife In­ter­na­tional, 2009; BirdLife In­ter­na­tional, 2011)

Tufted ducks are not en­dan­gered. They live in a large area and their num­bers are not de­creas­ing. On the IUCN Red list, they are listed as a species of "least con­cern". Their biggest threat is loss of habi­tat due caused by human ac­tiv­i­ties, in­clud­ing de­stroy­ing wet­lands to build build­ings, and pol­lu­tion from trash or oil spills. (BirdLife In­ter­na­tional, 2009; BirdLife In­ter­na­tional, 2011)

Con­trib­u­tors

MayaV. Azzi (au­thor), Rad­ford Uni­ver­sity, RyanJ Gar­ri­son (au­thor), Rad­ford Uni­ver­sity, Chris­tine Small (ed­i­tor), Rad­ford Uni­ver­sity, Rachelle Ster­ling (ed­i­tor), Spe­cial Pro­jects, Cather­ine Kent (ed­i­tor), Spe­cial Pro­jects.

Glossary

Ethiopian

living in sub-Saharan Africa (south of 30 degrees north) and Madagascar.

World Map

Nearctic

living in the Nearctic biogeographic province, the northern part of the New World. This includes Greenland, the Canadian Arctic islands, and all of the North American as far south as the highlands of central Mexico.

World Map

Pacific Ocean

body of water between the southern ocean (above 60 degrees south latitude), Australia, Asia, and the western hemisphere. This is the world's largest ocean, covering about 28% of the world's surface.

World Map

Palearctic

living in the northern part of the Old World. In otherwords, Europe and Asia and northern Africa.

World Map

acoustic

uses sound to communicate

agricultural

living in landscapes dominated by human agriculture.

bilateral symmetry

having body symmetry such that the animal can be divided in one plane into two mirror-image halves. Animals with bilateral symmetry have dorsal and ventral sides, as well as anterior and posterior ends. Synapomorphy of the Bilateria.

chemical

uses smells or other chemicals to communicate

coastal

the nearshore aquatic habitats near a coast, or shoreline.

colonial

used loosely to describe any group of organisms living together or in close proximity to each other - for example nesting shorebirds that live in large colonies. More specifically refers to a group of organisms in which members act as specialized subunits (a continuous, modular society) - as in clonal organisms.

cryptic

having markings, coloration, shapes, or other features that cause an animal to be camouflaged in its natural environment; being difficult to see or otherwise detect.

diurnal
  1. active during the day, 2. lasting for one day.
endothermic

animals that use metabolically generated heat to regulate body temperature independently of ambient temperature. Endothermy is a synapomorphy of the Mammalia, although it may have arisen in a (now extinct) synapsid ancestor; the fossil record does not distinguish these possibilities. Convergent in birds.

estuarine

an area where a freshwater river meets the ocean and tidal influences result in fluctuations in salinity.

female parental care

parental care is carried out by females

food

A substance that provides both nutrients and energy to a living thing.

freshwater

mainly lives in water that is not salty.

iteroparous

offspring are produced in more than one group (litters, clutches, etc.) and across multiple seasons (or other periods hospitable to reproduction). Iteroparous animals must, by definition, survive over multiple seasons (or periodic condition changes).

marsh

marshes are wetland areas often dominated by grasses and reeds.

migratory

makes seasonal movements between breeding and wintering grounds

monogamous

Having one mate at a time.

motile

having the capacity to move from one place to another.

natatorial

specialized for swimming

native range

the area in which the animal is naturally found, the region in which it is endemic.

omnivore

an animal that mainly eats all kinds of things, including plants and animals

oviparous

reproduction in which eggs are released by the female; development of offspring occurs outside the mother's body.

saltwater or marine

mainly lives in oceans, seas, or other bodies of salt water.

seasonal breeding

breeding is confined to a particular season

sexual

reproduction that includes combining the genetic contribution of two individuals, a male and a female

social

associates with others of its species; forms social groups.

tactile

uses touch to communicate

temperate

that region of the Earth between 23.5 degrees North and 60 degrees North (between the Tropic of Cancer and the Arctic Circle) and between 23.5 degrees South and 60 degrees South (between the Tropic of Capricorn and the Antarctic Circle).

terrestrial

Living on the ground.

territorial

defends an area within the home range, occupied by a single animals or group of animals of the same species and held through overt defense, display, or advertisement

tropical savanna and grassland

A terrestrial biome. Savannas are grasslands with scattered individual trees that do not form a closed canopy. Extensive savannas are found in parts of subtropical and tropical Africa and South America, and in Australia.

savanna

A grassland with scattered trees or scattered clumps of trees, a type of community intermediate between grassland and forest. See also Tropical savanna and grassland biome.

temperate grassland

A terrestrial biome found in temperate latitudes (>23.5° N or S latitude). Vegetation is made up mostly of grasses, the height and species diversity of which depend largely on the amount of moisture available. Fire and grazing are important in the long-term maintenance of grasslands.

visual

uses sight to communicate

young precocial

young are relatively well-developed when born

Ref­er­ences

Bent, A. 1951. Life His­to­ries of North Amer­i­can Wild Fowl Ducks, Geese, and Swans. New York: Dover Pub­li­ca­tions Inc.

Bevan, R., J. Speak­man, P. But­ler. 1995. Daily en­ergy ex­pen­di­ture of tufted ducks: a com­pair­son be­tween in­di­rect calorime­try dou­bly la­beled water and heart rate. Func­tional Ecol­ogy, 9: 40-47.

BirdLife In­ter­na­tional, 2011. "Birdlife In­ter­na­tional" (On-line). Species fact­sheet: Aythya fuligula. Ac­cessed Feb­ru­ary 16, 2011 at http://​www.​birdlife.​org/​datazone/​speciesfactsheet.​php?​id=480.

BirdLife In­ter­na­tional, 2009. "The IUCN Red List of Threat­ened Species" (On-line). Ac­cessed March 21, 2011 at www.​iucnredlist.​org.

Blums, P., J. Nichols, M. Lind­berg, J. Hines, A. Med­nis. 2003. Fac­tors af­fect­ing breed­ing dis­per­sal of Eu­ro­pean ducks on Eu­gene Marsh, Latvia. Jour­nal of An­i­mal Ecol­ogy, 72: 292-307.

Cleeves, T. 2002. RSPB Hand­book of British Birds. Lon­don: A & C Black Pub­lish­ers Ltd.

De Leeuw, J. 1999. Food in­take rates and habi­tat seg­re­ga­tion of tufted duck Aythya fuligula and scaup Aythya mar­ila ex­ploit­ing zebra mus­sles Dreis­sena poly­mor­pha. Ardea, 87: 15-31.

Dick, G. 2002. "Field Guide to Birds of North Amer­ica" (On-line). Tufted Duck. Ac­cessed Feb­ru­ary 15, 2011 at http://​identify.​whatbird.​com/​obj/​420/_/​Tufted_​Duck.​aspx.

Draulans, D. 1982. For­ag­ing and size se­lec­tion of mus­sles by the tufted duck Aythya fuligula. Jour­nal of An­i­mal Ecol­ogy, 51: 943-956.

Green, J., L. Halsey, P. But­ler. 2005. To what ex­tent is for­ag­ing be­hav­ior of aquatic birds con­strained by their phys­i­ol­ogy?. Phys­i­o­log­i­cal and Bio­chem­i­cal Zo­ol­ogy, 78, 5: 766-781.

Halsey, L., S. Wal­lace, A. Woakes, H. Win­kler, P. But­ler. 2005. Tufted ducks Aytha fuligula do not con­trol buoy­ancy dur­ing div­ing. Jour­nal of Avian Bi­ol­ogy, 36: 261-267.

Hill, D. 1983. Lay­ing date, clutch size and eggs size of the mal­lard Anas platyrhyn­chos and tufted duck Aythya fuligula. IBIS, 126: 484-495.

Hill, D. 1984. Fac­tors af­fect­ing nest suc­cess in the mal­lard and tufted duck. Ornis Scan­di­nav­ica, 15/2: 115-122.

Mason, B. 1996. A Lit­tle Oasis: The Early His­tory of Ash­ton Park West Kirby. 1 & 3 Grove Road, Rock Ferry, Birken­head Wirral CH42 3XS: Coun­tyvise Ltd.

Miller, G., S. Spool­man. 2008. Liv­ing in the En­vi­ron­ment: Prinic­ples, Con­nec­tions, and So­lu­tions. Flo­rence, Ken­tucky: Cen­gage Learn­ing, Inc.

Nils­son, L. 1972. Habi­tat se­lec­tion, food choice, and feed­ing bai­tats of div­ing ducks in coastal wa­ters of South Swe­den dur­ing non-breed­ing sea­son. Ornis Scan­di­nav­ica, 3: 55-78.

Nils­son, L. 2005. Long-term trends and changes in num­bers and dis­tri­b­u­tion of some win­ter­ing wa­ter­fowl species. Acta Zo­o­log­ica Litu­anica, 15/2: 151-157.

Ny­s­trom, K., O. Pehrs­son. 1988. Salin­ity as a con­straint af­fect­ing food and habi­tat choice of mus­cle-feed­ing div­ing ducks. IBIS, 130: 94-110.

Oka, N., M. Ya­ma­muro, J. Hiart­suka, H. Satoh. 1999. Habi­tat se­lec­tion by win­ter­ing tufted ducks with spe­cial ref­er­ence to their di­ges­tive organ and to pos­si­ble seg­re­ga­tion be­tween neigh­bor­ing pop­u­la­tions. Eco­log­i­cal Re­search, 14: 303-315.

Olney, P. 1963. The food and feed­ing habits of tufted duck Aythya fuligula. IBIS, 105/1: 55.

Robin­son, R. 2005. "Tufted Duck Aythya fuligula" (On-line). BTO look­ing out for birds. Ac­cessed March 17, 2011 at http://​blx1.​bto.​org/​birdfacts/​results/​bob2030.​htm.

Rudol­fova, J., J. Sitko, P. Horak. 2002. Nasal schis­to­somes of wild­fowl in the Czech Re­pub­lic and Poland. Folia Par­a­sito­log­ica, 88: 1093-1095.

The Wild­fowl Trust, , Slim­bridge, Glos. 1963. Food and feed­ing habits of the tufted duck Aythya fuligu­lia. In­ter­na­tional Jour­nal of Avi­a­tion and Sci­ence, 105;1: 55-62.

Win­field, I., D. Win­field. 1994. Feed­ing ecol­ogy of the div­ing ducks por­chard (Aythya feri­nal), tufted duck (A. fuligula), scarp (A. mar­ila) and gold­en­eye (Buchep­hala clan­gula) over win­ter­ing on Lough Neagh, North­ern Ire­land. Fresh­wa­ter Bi­ol­ogy, 32: 467-477.