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Home -> Kingdom Animalia -> Phylum Chordata -> Subphylum Vertebrata -> Class Aves -> Order Passeriformes -> Family Icteridae -> Species Sturnella magna

Sturnella magna
eastern meadowlark



2009/11/22 05:04:54.101 US/Eastern

By Tamar Dexheimer

Kingdom: Animalia
Phylum: Chordata
Subphylum: Vertebrata
Class: Aves
Order: Passeriformes
Family: Icteridae
Genus: Sturnella
Species: Sturnella magna

Geographic Range

Sturnella magna is found in the eastern United States, as well as parts of the southwest U.S. and Central America. The summer breeding range includes parts of southern Canada. ("Eastern Meadowlark", 1992; Campbell, 1973)

Biogeographic Regions:
nearctic (native ); neotropical (native ).

Habitat

Eastern meadowlarks breed in native grasslands, pastures, savannas, alfalfa and hay fields, cropland borders, roadsides, orchards, golf courses, airports, reclaimed strip mines, overgrown fields, and other open areas. In the western range, the breeding range also consists of tall-grass prairies and desert grassland. In the winter they are generally found in open country, cultivated fields, feedlots, and marshes. Eastern meadowlarks are generally found in habitats that are more mesic than their close relative, western meadowlarks (S. neglecta). ("Eastern Meadowlark", 1992; Elliott and Read, 1998; Lanyon, 1995)

These animals are found in the following types of habitat:
temperate ; tropical ; terrestrial .

Terrestrial Biomes:
savanna or grassland .

Physical Description

Mass
90 to 150 g; avg. 120 g
(3.17 to 5.28 oz; avg. 4.22 oz)


Length
19 to 26 cm
(7.48 to 10.24 in)


Wingspan
35 to 40 cm
(13.78 to 15.75 in)


Eastern meadowlarks are medium-sized songbirds, with long, slender, light gray bills and dark brown eyes. The tails are short and have rigid rectrices. The legs and toes are long. Male S. magna have grayish heads with blackish stripes, a yellow “eyebrow”, and dark crowns with a median stripe. The wings and tail are streaked and barred with dark and light brown. Males have a broad white moustachial stripe and a yellow chin, which is divided from the underparts by a broad black breast band. The underparts turn off-white on the streaked flanks and under the tail coverts. The pale undertail coverts are streaked and spotted dusky black. Females are similar to males except that they are smaller, paler, and have a narrower breast band. Males are slightly larger than females, from 21 to 25 cm in length, females are from 19 to 23 cm. Juvenile eastern meadowlarks have masked black areas and the white areas are buffish. Juveniles also have more brown plumage in the winter. Eastern meadowlark eggs are white, speckled with reddish-brown. When these birds walk, the tail constantly jerks open. These birds fly by beating their wings vigorously and then gliding. ("Eastern Meadowlark", 1992; Birdnature.com, 2002; Campbell, 1973; Lanyon, 1995)

Some key physical features:
endothermic ; homoiothermic; bilateral symmetry .

Sexual dimorphism: male larger, sexes colored or patterned differently.

Reproduction

Breeding interval
Breeding first occurs in late May, with a second brood produced in late June to early July.

Breeding season
Breeding occurs from late May to August.

Eggs per season
6 to 14; avg. 10

Time to hatching
13 to 15 days

Time to fledging
11 to 12 days

Time to independence
2 weeks (low)

Age at sexual or reproductive maturity (female)
1 years (average)

Age at sexual or reproductive maturity (male)
1 years (average)

Male eastern meadowlarks are polygynous, with most males having two to three mates. Female S. magna have only one mate per breeding season, provided that the male successfully defends the territory. Males establish their territories approximately two to four weeks before females arrive. Male S. magna display their territories with flight displays and by singing. Female eastern meadowlarks choose their mates by selecting territories, which are defended by males with conspecific vocalizations. Once the pair bond forms the pair remains close together while foraging and searching for nest sites. A male S. magna defends its territory against rivals by fluffing out its plumage and pointing its bill upwards. Males guard their mates from neighboring males by constantly guarding their mate. (Campbell, 1973; Cornell Lab of Ornithology, 2005a; Lanyon, 1995)

Males establish their territories in March, females arrive about two to four weeks later females. Male eastern meadowlarks rarely engage in body contact and fighting when defending their territories, however, when it does occur it can be quite severe. Pairing occurs immediately after females arrive. The "aerial chase" occurs within minutes of a female choosing a male. The female typically initiates the chase, although sometimes the chase includes two females and one male. The aerial chase consists of either a series of short flights or as brief flights interspersed with periods of posturing and rest. Additionally, the male is typically silent during the aerial chase. These chases usually carry the participants well beyond the boundaries of the male’s territory. When a female eastern meadowlark is receptive, she eventually assumes the receptive posture, at which time the male will approach, paw the female’s back and then mount. Afterwards the female remains in a semi-receptive position and flutters and shakes its plumage, chatters several times, then vigorously preens itself. The female receptive posture consists of the female elevating its bill and tail, holding its wings slightly drooped, and quivering, sometimes the female also chatters. Later on in the breeding season "jump-flights" and tee-tee-tee calls may accompany the receptive posture. However, if a male approaches when the female is not receptive, the female will use "expansion posturing" to warn off the male. Also, males and females make jump-flights before and during repeated copulation periods. A jump-flight consists of the bird jumping approximately one meter into the air and then flying several meters. Once the breeding season is over, male S. magna cease defending their territories. (Campbell, 1973; Cornell Lab of Ornithology, 2005a; Francq, 1972; Lanyon, 1995)

Mating systems:
polygynous .

Female eastern meadowlarks gather nest materials and build the nest. The nest consists of coarse grasses, lined with finer grasses and is constructed on the ground, typically in a shallow depression. The outside diameter of the nest ranges from 14-21 cm, the inside diameter ranges from 8-15 cm, and the inside depth ranges from 5-8 cm. Female S. magna land a distance away from the nest and then stealthily approach the nest. (Campbell, 1973; Cornell Lab of Ornithology, 2005a; Lanyon, 1995)

Key reproductive features:
iteroparous ; seasonal breeding ; gonochoric/gonochoristic/dioecious (sexes separate); sexual ; oviparous .

Females incubate the eggs for 13 to 15 days, when the altricial young hatch. After the eggs hatch both the female and her mate feed the hatchlings. However, females do most of the feeding. Nestlings typically fledge 11 to 12 days after hatching, but juveniles do not become independent for at least another two weeks. The parents continue to feed the fledglings until they become independent. (Campbell, 1973; Cornell Lab of Ornithology, 2005b; Lanyon, 1995)

Parental investment:
altricial ; pre-fertilization (provisioning, protecting: female); pre-hatching/birth (provisioning: female, protecting: female); pre-weaning/fledging (provisioning: male, female, protecting: female); pre-independence (provisioning: male, female, protecting: female).

Lifespan/Longevity

Extreme lifespan (wild)
9 years (high)

Average lifespan (wild)
5 years

Typical lifespan (captivity)


Eastern meadowlarks have an expected lifespan of five years in the wild, which is the same as the high end of its expected lifespan in captivity. The longest know lifespan in the wild is nine years. (Lanyon, 1995)

Behavior

Territory Size
28 to 32 m^2

Eastern meadowlarks are social, forming loose flocks during the fall and winter. These flocks lack a social hierarchy and are simply a loose aggregation of S. magna and occasionally, S. neglecta (western meadowlarks). They use a variety of songs, calls and postures to communicate with other meadowlarks. Also, where eastern and western meadowlark ranges overlap, male eastern meadowlarks will defend against male eastern and western meadowlarks. Males typically defend their territories with posturing and aerial displays.

Both male and female S. magna often preen and stretch, especially in the early morning hours. Stretching, specifically of the legs and wings, usually follows preening. They also tend to scratch their head with their foot, which they bring up over their wing. Sturnella magna bathes in puddles and wet grass. Sturnella magna roosts on the ground in thick grass, with its head under its scapulars and its body resting on the ground. (Cornell Lab of Ornithology, 2005a; Lanyon, 1995)

Sturnella magna do not migrate, except for those in the northernmost parts of their range. However, S. magna tend to form flocks during the winter. Also, those S. magna that do migrate, do so during daylight hours, and begin their migration when it begins to freeze and snow. Some of these birds may migrate over 1,000 km to their winter range. Eastern meadowlarks that migrate, leave by the end of November and return to the breeding range in March. (Francq, 1972; Lanyon, 1995)

Home Range

Male S. magna establish their territories in March, and defend their territories throughout the breeding season. During the breeding season the territories change in size and shape depending on population densities, relocations of female activity centers, and changes in habitat suitability. Once the breeding season is over S. magna do not defend or maintain territories. (Francq, 1972; Lanyon, 1995)

Key behaviors:
terricolous; flies; diurnal ; motile ; migratory ; sedentary ; territorial ; social .

Communication and Perception

The songs of S. magna are one of the first birdsongs of spring. Sturnella magna have a variety of vocal communications. There are begging notes, location notes, dzert, whistle, chatter, weet, primary song, flight song, female song, zeree, and tee-tee-tee. Nestlings and recently fledged juveniles use begging and location notes, which are simple high-pitched notes. These notes enable the parents to find and feed their young. The dzert call indicates mild disturbance. The whistle indicates intense excitement in males or females, such as the presence of a predator, just before a flight song, or immediately after an aerial chase or copulation. Both sexes use the chatter call to indicate excitement such as the presence of a predator or intruder. Females also chatter after copulation and in response to their mates’ primary song. Only males use the primary song, which sounds like seee-yeee, seee-yer. In the courtship period, female S. magna use the female song, during early morning preening. The alarm call of the eastern meadowlark is a short buzzy, dzert. ("Eastern Meadowlark", 1992; Cornell Lab of Ornithology, 2005a; Elliott and Read, 1998; Lanyon, 1995)

Posturing and aerial chases are used to attract and pursue possible mates. Jump-flights are used to ward off males that are intruding on another male’s territory. Bill-tilting and tail- and wing-flashing are used in territorial disputes, as is expansion posturing. Expansion posturing is when individuals extend their contour feathers, spread the tail, and draws the head close to the body. Female S. magna use expansion posturing to warn off its mate when the female is unreceptive. If expansion posturing does not succeed in warning off the male, the female will hold its feathers tight against its body and point its gaping bill at the male. Male eastern meadowlarks also use expansion posturing after the formation of the pair bond. ("Eastern Meadowlark", 1992; Cornell Lab of Ornithology, 2005a; Elliott and Read, 1998; Lanyon, 1995)

Communicates with:
visual ; acoustic .

Perception channels:
visual ; tactile ; acoustic ; chemical .

Food Habits

Eastern meadowlarks walk and run on the ground while foraging for food, they also forage by probing beneath the soil. Their diet varies with the season. In the spring they feed mainly on cutworms, grubs, and caterpillars. When summer comes they eat insects, primarily beetles and grasshoppers. In the winter they eat noxious weed seeds and waste grains as well as some wildfruits and occasional carrion from road-kill or predator-kills. (Campbell, 1973; Lanyon, 1995)

Animal Foods:
carrion ; insects.

Plant Foods:
seeds, grains, and nuts; fruit.

Predation

Known predators

Eastern meadowlarks are preyed on by hawks and falcons and occasionally by owls. They are most likely to be preyed upon by owls during the owl’s breeding season. While the owls are raising their young, they are more likely to hunt during daylight hours, in order to catch enough prey to feed the chicks. Hawks and falcons are diurnal, and often hunt in similar habitats. During their nesting season, domestic cats, dogs, foxes, coyotes, and skunks prey upon the eggs and nestlings. Eastern meadowlark coloration helps them to blend in to their grassland surroundings, they can be difficult to spot unless they are on a high perch. (Grossman and Hamlet, 1964; Lanyon, 1995)

Anti-predator adaptations::
cryptic .

Ecosystem Roles

Eastern meadowlarks are prey for larger predators and they prey on a variety of insects, including grubs and caterpillars, which could damage the surrounding vegetation. They also act to disperse the sees of plants they eat. Sturnella magna serves as a host for a variety of internal and external parasites, and for brown-headed cowbirds. Brown-headed cowbirds are obligate parasites, which lay eggs in the nests of other species of birds. ("Western Meadowlark", 2003; Birdnature.com, 2002; Campbell, 1973; Cornell Lab of Ornithology, 2005c; Grossman and Hamlet, 1964; Lanyon, 1995; Stark, 1940; Taylor, 1969)

Key ways these animals impact their ecosystem:
disperses seeds; biodegradation ; soil aeration .

Commensal or parasitic species (or larger taxonomic groups) that use this species as a host

Economic Importance for Humans: Negative

Sturnella species eat kernels of sprouting grain, which can destroy portions of newly planted crops. ("Western Meadowlark", 2003)

Ways that these animals might be a problem for humans:
crop pest.

Economic Importance for Humans: Positive

Sturnella magna eat insects that are crop pests, therefore they act to control pest populations that impact crops. (Campbell, 1973; Lanyon, 1995)

Ways that people benefit from these animals:
controls pest population.

Conservation Status

IUCN Red List: [link]:
Least Concern.

US Migratory Bird Act: [link]:
No special status.

US Federal List: [link]:
No special status.

CITES: [link]:
No special status.

According to the IUCN Red List, the U.S. Federal List, and the State of Michigan List, eastern meadowlarks have no special status. They are not threatened, likely to become threatened, or endangered. This agrees with the Audubon Society's assessment of S. magna. Eastern meadowlarks fall into the Audubon Society's green conservation status, which means that it is of low or no conservation concern. However, S. magna populations have been experiencing a significant population decline, declining by as much as 50% since 1966. (National Audubon Society, 2005)

Other Comments

Eastern meadowlarks are not true larks; rather they belong to the same family as blackbirds and orioles (Icteridae). There are about 18 recognized subspecies of the eastern meadowlark.

The decline of the S. magna populations could be partially due to the industrialization of agriculture, which increases the likelihood of a nest being destroyed by the agricultural machinery and the increased use of row crops which are an unsuitable habitat for these birds. Another possible cause of the decline is apparent predation by cattle. Cattle have been documented destroying nests, sometimes by accident but also by crushing eggs and nestlings with their muzzles and by removing nestlings from the nests. (Cornell Lab of Ornithology, 2005c; Nack and Ribic, 2005)

For More Information

Contributors

Tanya Dewey (editor), Animal Diversity Web, University of Michigan Museum of Zoology.

Tamar Dexheimer (author), Kalamazoo College. Ann Fraser (editor, instructor), Kalamazoo College.

References

1992. Eastern Meadowlark. Pp. 345 in R. O. Zeleny, ed. The World Book Encyclopedia, Vol. 13. Chicago: World Book Inc..

2003. Western Meadowlark. Pp. 316 in M. Hutchins, ed. Grzimek's Animal Life Encyclopedia, Vol. 11. Detroit: Gale Group Inc..

Birdnature.com. 2002. "Birdnature.com" (On-line). Accessed October 01, 2005 at http://www.birdnature.com/meadowlark.html.

Campbell, B. 1973. Sturnella magna. Pp. 337 in R. T. Holmes, ed. The Dictionary of Birds in Color. New York: The Viking Press.

Cornell Lab of Ornithology. 2003. "Eastern Meadowlark" (On-line). Accessed November 13, 2005 at http://www.birds.cornell.edu/programs/AllAboutBirds/BirdGuide/Eastern_Meadowlark.html.

Cornell Lab of Ornithology. 2005. "Behavior" (On-line). Birds of North America Online. Accessed November 19, 2005 at http://bna.birds.cornell.edu/BNA/account/Eastern_Meadowlark/BEHAVIOR.html.

Cornell Lab of Ornithology. 2005. "Breeding" (On-line). Birds of North America Online. Accessed November 19, 2005 at http://bna.birds.cornell.edu/BNA/account/Eastern_Meadowlark/BREEDING.html.

Cornell Lab of Ornithology. 2005. "Demography and Populations" (On-line). Birds of North America Online. Accessed November 19, 2005 at http://bna.birds.cornell.edu/BNA/account/Eastern_Meadowlark/DEMOGRAPHY_AND_POPULATIONS.html.

Elliott, L., M. Read. 1998. Common Birds And Their Tongs. Boston: Houghton Mifflin Company.

Francq, G. 1972. Parental care of the eastern meadowlark (Sturnella magna). Kansas: Kansas State Teachers College.

Grossman, M., J. Hamlet. 1964. Birds of Prey of the World. New York: Bonanaza Books.

Lanyon, W. 1995. Eastern Meadowlark: Sturnella magna. Washington D. C.: American Ornithologists' Union.

Nack, J., C. Ribic. 2005. Apparent predation by cattle at grassland bird nests. Wilson Bulletin, 117: 56-62.

National Audubon Society. 2005. "State of the Birds: Grasslands" (On-line). Accessed October 11, 2005 at http://www.audubon.org/bird/stateofthebirds/grasslands.html.

Stark, F. 1940. A study of the animal parasites of Sturnella magna magna and Sturnella neglecta of southeastern Kansas. Pittsburg, Kansas: Kansas State Teachers College.

Taylor, R. 1969. Histological study of host-parasite relations between meadowlarks (Sturnella) and Microtetrameres Sturnellae (Nematoda: Tetrameridae). Oklahoma: University of Oklahoma Graduate College.

2009/11/22 05:04:57.076 US/Eastern

To cite this page: Dexheimer, T. and A. Fraser. 2006. "Sturnella magna" (On-line), Animal Diversity Web. Accessed November 23, 2009 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Sturnella_magna.html.

Disclaimer: The Animal Diversity Web is an educational resource written largely by and for college students. ADW doesn't cover all species in the world, nor does it include all the latest scientific information about organisms we describe. Though we edit our accounts for accuracy, we cannot guarantee all information in those accounts. While ADW staff and contributors provide references to books and websites that we believe are reputable, we cannot necessarily endorse the contents of references beyond our control.

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