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Spheniscus humboldti
Humboldt penguin


By Jillian Smith

Kingdom: Animalia
Phylum: Chordata
Subphylum: Vertebrata
Class: Aves
Order: Sphenisciformes
Family: Spheniscidae
Genus: Spheniscus
Species: Spheniscus humboldti

Geographic Range

Spheniscus humboldti inhabit the coastal regions of Peru and Chile. These regions are temperate in climate (Welch 1994). These birds are well known in the Humboldt current from Peru to south of Chile on the coast and offshore islands (Villouta, et al., 1997).

Biogeographic Regions
neotropical (Native )

Habitat

Range depth
1000 (high) m
(3280.84 (high) ft)

On land Spheniscus humboldti lives in burrows composed of soil and rock (Welch 1994). Spheniscus humboldti breed in large colonies. They spend most of their time at sea and rarely come back to land (Chicago Zoological Society 1999).

Habitat Regions
temperate

Aquatic Biomes
coastal

Physical Description

Average mass
4000 g
(140.97 oz)

Range length
38 (low) cm
(14.96 (low) in)

Spheniscus humboldti are black and white in color with pink around the eyes and on the beak. The feet are webbed and serve as a rudder. There are also claws at the end of the toes for climbing. The feathers are in two layers. The top layer is flat and overlaps the second layer to stop the wind and water from penetrating to the body. The second down layer is for insulation. The wings evolved into flippers for flying through the water. The bones are solid and act as a ballast while diving (The Smithsonian Zoo 2001). The body is in the shape of a streamlined torpedo covered by the short waterproof feathers (Chicago Zoological Society 1999). They are able to swim swiftly through the water by the aid of hard flippers or wings (The Aquatic Creatures 2002). Spheniscus humboldti is also called the Peruvian penguin. It is 38 to 45 centimeters (18 to 15 inches) in height and weighs about 4 kg (9 pounds) (The Smithsonian Zoo 2001).

Other Physical Features
endothermic ; bilateral symmetry

Reproduction

One male breeds with one female during the mating season (Busch Entertainment Corp. 2000).

Mating System
monogamous

Breeding season
year round

Average eggs per season
2

Average time to hatching
39 days

Average age at sexual or reproductive maturity (female)
3 years

Average age at sexual or reproductive maturity (male)
3 years

The most abundant breeding happens around the availability of food and of nesting sites. Egg laying occurs throughout the year (Welch 1994). Once a mate is acquired the pair initiates copulations after displays of behavior (The Smithsonian Zoo 2001). Spheniscus humboldti begin breeding at about three years of age. The male arrives at the nesting area a few days before the female to prepare the nesting site. The female arrives and lays two white eggs that she incubates for 39 days (Chicago Zoological Society 1999). The major causes of egg loss are from flooding of nests during ocean storms, accidental breakage, and nest desertion and predation by gulls (International Conservation Work Group 2001). Chicks poke a small hole through the egg then chip the shell off. This can take up to three days. (Busch Entertainment Corp. 2000).

Key Reproductive Features
iteroparous ; year-round breeding ; gonochoric/gonochoristic/dioecious (sexes separate); sexual ; oviparous

Chicks require attentive parents. They depend on the parents for survival between hatching and growing waterproof feathers. Once a chick has done this it can enter the water and become independent (Busch Entertainment Corp. 2000).

Parental Investment
altricial ; male parental care ; female parental care

Lifespan/Longevity

The average life span is 15 to 20 years for Spheniscus humboldti and there is a high mortality rate among the young (Busch Entertainment Corp. 2000).

Behavior

The feathers must be groomed to prevent water infiltration as maintaining a high degree of air insulation is important. Before their early feeding in the morning, Spheniscus humboldti preen. This requires gathering oil from the preening gland located in the rump where the tail originates. The oil is applied to the feathers and edges of the flippers. They will also preen each other. A lot of huddling is done to minimize heat loss. Spheniscus humboldti, however, can get hot through physical activities. Stream-lining feathers reduce the volume of air between the skin and the edge of the feathers and reduces the layer of air that serves as insulation. Molting takes place sometime in July to September and it only last about ten days (The Smithsonian Zoo 2001).

Key Behaviors
social ; colonial

Food Habits

Spheniscus humbldti are inshore feeders; they forage for small fish and crustaceans. They circle around the prey and attack from the side swallowing it head first (Welch 1994). The mouth and tongue have backward pointed spines to hold fish (The Smithsonian Zoo 2001). El Nino storms destroy large regions of nesting areas by causing rough surf that washes away nests. The affects on the temperature of the sea is an increase which can reduce the food supply (Chicago Zoological Society 1999).

Primary Diet
carnivore (Piscivore , Eats non-insect arthropods)

Animal Foods
fish; aquatic crustaceans

Predation

Known Predators


Spheniscus humboldti can swim up to 30 miles per hour and are very agile when swimming. This is their only defense against predators (The Smithsonian Zoo 2001). When in the water they can be eaten by leopard seals, fur seals, sea lions, sharks, and killer whales. On land foxes, snakes, and introduced predators like cats and dogs can prey on the eggs and chicks (Busch Entertainment Corp. 2000). Spheniscus humboldti get entangled in fishing nets as well (Chicago Zoological Society 1999).

Ecosystem Roles

Spheniscus humboldti live on the coast and gather soil, rocks, and sometimes grasses, to build their nests. These nests are created using their wings and feet to push and mold a nest (Welch 1994).

Ecosystem Impact
creates habitat

Economic Importance for Humans: Positive

Humans collect the eggs of Spheniscus humboldti (Chicago Zoological Society 1999). In the nineteenth century penguin skins were used to make caps, slippers, and purses. The feathers were used for clothing decorations. Extraction of oil from the penguins fat layers was economically important; the oil was used for lighting, tanning leather, and fuel. Spheniscus humboldti guano had a value as nitrogen rich fertilizer (Busch Entertainment Corp. 2000).

Positive Impacts
produces fertilizer

Conservation Status

IUCN Red List of Threatened Species [Link]
Vulnerable
More Information

US Migratory Bird Act [Link]
Protected

US Federal List [Link]
Endangered

CITES [Link]
Appendix I

In 1981 the U.S. department of Interior declared Spheniscus humboldti endangered. Today Spheniscus humboldti are only used illegally (Welch 1994).

For More Information

Find Spheniscus humboldti information at

Contributors

Jillian Smith (author), Western Maryland College, Randall L. Morrison (editor), Western Maryland College.

References

Buda, J., N. Parikh, J. Kim. 2001. "The Humboldt Penguin: *Spheniscus humboldti*" (On-line). Accessed March 8, 2002 at http://www.bergen.org/Smithsonian/HumboldtPenguin/.

Busch Entertainment Corp., 2000. "Penguins" (On-line). Accessed April 21, 2002. at http://www.seaworld.org/infobooks/Penguins/home.html.

Chicago Zoological Society, 1999. "Fact Sheet: Humboldt Penguin" (On-line). Accessed March 1, 2002 at http://www.letus.org/bmatters/animals/penguin.html.

International Conservation Work Group, 2001. "Humboldt Penguin" (On-line). Accessed March 21, 2002 at http://www.penguins.ac/humboldt-penguins.htm.

The Aquatic Creatures, 2002. "Peruvian Penguin:Swimming" (On-line). Accessed March 18, 2002 at http://www3.ocn.ne.jp/~kmitoh/zaturoku/penwing/e_penwing.html.

Villouta, G., R. Hargreaves, V. Riveros. 1997. Haematological and clinical biochemistry findings in captive Humboldt penguins (Spheniscus humboldti). Avian Pathology, 26: 851-858.

Welch, K. 1994. "The Penguin Page: A Journey to the Bottom of the World" (On-line). Accessed March 15, 2002 at http://users.capu.net/~kwelch/pp/species/humboldt.html.

To cite this page: Smith, J. 2003. "Spheniscus humboldti" (On-line), Animal Diversity Web. Accessed June 01, 2012 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Spheniscus_humboldti.html

Disclaimer: The Animal Diversity Web is an educational resource written largely by and for college students. ADW doesn't cover all species in the world, nor does it include all the latest scientific information about organisms we describe. Though we edit our accounts for accuracy, we cannot guarantee all information in those accounts. While ADW staff and contributors provide references to books and websites that we believe are reputable, we cannot necessarily endorse the contents of references beyond our control.

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Structured Inquiry Search — preview