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Pholcus phalangioides


By Anna Ferrick

Kingdom: Animalia
Class: Arachnida
Order: Araneae
Family: Pholcidae
Genus: Pholcus
Species: Pholcus phalangioides

Geographic Range

Pholcus phalangioides is found throughout the world. It is a common cellar spider throughout the United States. (Emerton, 1902; Jackman, 1997)

Other Geographic Terms
cosmopolitan

Habitat

Pholcus phalangiodes can be found in undisturbed, low light locations. Some places one might encounter this spider are in basements, under stones, under ledges, and in caves. People most often associate these spiders with living on ceilings and in corners in homes. They make their webs large, loose, and flat, but they can make them in irregular shapes to fit into surrounding objects. Their webs are normally oriented horizontally. Pholcus phalangioides hangs upside down on the web it makes. (Emerton, 1902; Foelix, 1982; Jackman, 1997; Uhl, November 1998)

Habitat Regions
temperate ; tropical ; terrestrial

Terrestrial Biomes
savanna or grassland ; forest

Other Habitat Features
urban ; suburban ; agricultural ; caves

Physical Description

Range length
6 to 8 mm
(0.24 to 0.31 in)

Pholcus phalangioides is pale yellow-brown except for a large gray patch in the center of the cephalothorax. The body and legs are almost translucent. These spiders are covered with fine gray hairs. The head is a darker color around the eyes. A translucent line marks the dorsal vessel. There are eight eyes: two small eyes in front of the two triads of larger eyes.

Females are seven to eight millimeters in length and males are six millimeters.

Because of the translucent quality of this animal, using a microscope it is possible to see the moving blood cells in the legs and body of a living animal. (Jackman, 1997)

Other Physical Features
bilateral symmetry

Sexual Dimorphism
female larger

Development

A mother P. phalangioides watches over her newly hatched young (prenymphs) for about nine days until the prenymphs shed their skins to become little spiders. The young spiders then leave the maternal web, and go look for a place to build their own webs. (Stüber, 1999)

Reproduction

In studies done by Gabriele Uhl at the University of Bonn, male P. phalangioides seemed to be attracted to and to mate with larger females more often than smaller females. This may increase reproductive success for males, because large females produce more eggs than smaller females. (Uhl, November 1998)

Breeding season
Peak breeding occurs between June and September.

Before mating, a male spider deposits some sperm onto a little web, and then sucks it into a special cavity within his pedipalp. During mating, which can take several hours, the male deposits his sperm into the female's epigynum, which is an opening on the underside of her abdomen. Females can store the sperm in a special cavity at the beginning of the uterus, called the uterus externus, until it is time for her eggs to be fertilized. Timing of fertilization and laying depends on the availability of food. Because the sperm are stored for some period of time, it is possible for a female to mate again. If this occurs, the sperm from the two males mixes in the uterus externa. However, the sperm of the last male mated with has priority in fertilizing the eggs. This is because of a mechanism of sperm removal during mating. Males perform rhythmic movements of their intromittent organs during copulation, which results in extrusion of most of the sperm already in the uterus externa.

After a female lays her eggs, she wraps them in silk strands and carries the package in her chelicera (jaws), located on the underside of her body. (Emerton, 1902; Jackman, 1997; Stüber, 1999; Uhl, November 1998)

Key Reproductive Features
sexual ; fertilization (Internal ); sperm-storing

The only parental care female P. phalangioides offer their young is nine days of protection as the prenymphs finish developing into spiders. (Stüber, 1999)

Parental Investment
pre-fertilization (Provisioning, Protecting: Female); pre-hatching/birth (Provisioning: Female, Protecting: Female); pre-weaning/fledging (Protecting: Female)

Lifespan/Longevity

Range lifespan
Status: wild

3 (high) years

Range lifespan
Status: captivity

3 (high) years

Pholcus phalangioides can live up to about three years. (Stüber, 1999)

Behavior

When P. phalangioides is not mating, it is a solitary creature that work on catching food in its web. (Uhl, November 1998)

Key Behaviors
terricolous; troglophilic; motile ; solitary

Communication and Perception

Male spiders find a female spiders by tracking the pheromones she leaves. During mating, tactile communication is probably of some importance. (Jackman, 1997)

Communication Channels
tactile ; chemical

Other Communication Modes
pheromones

Perception Channels
visual ; tactile ; chemical

Food Habits

Pholcus phalangioides seems to prefer other spiders and small insects as prey. Also, males and females have both been known to engage in cannibalism. Females have been seen invading another spider's web, eating that spider, and using the foreign web to catch new prey for themselves. These spiders kill and digest their prey using venom. (Jackman, 1997; Stüber, 1999; Uhl, November 1998)

Animal Foods
insects; terrestrial non-insect arthropods

Predation

Known Predators
  • Birds


When the web of P. phalangioides is disturbed, the spider swings its body around rapidly with its legs attached firmly to its web. It swings fast enough that the spider becomes very hard to see. This may be a form of camouflage. (Emerton, 1902; Stüber, 1999)

Anti-predator Adaptations
cryptic

Ecosystem Roles

Because their diet is primarily insects, these spiders play the important role of controlling the growth of insect populations. (Stüber, 1999)

Economic Importance for Humans: Positive

Pholcus phalangioides helps control pest populations. (Uhl, November 1998)

Positive Impacts
controls pest population

Economic Importance for Humans: Negative

These spiders often live inside houses, where humans do not normally want spiders and their webs. (Uhl, November 1998)

Negative Impacts
household pest

Conservation Status

IUCN Red List [Link]
No special status

US Federal List [Link]
No special status

CITES [Link]
No special status

State of Michigan List [Link]
No special status

At this time Pholcus phalangioides are not endangered or threatened, so there is no conservation underway (ESA).

Other Comments

Because one of the common names for Pholcus phalangioides is daddylongleg spider, it is often confused with the Harvestmen daddylong legs, which are a separate species that is not even a true spider. Also, despite the urban myth that the daddylongleg spider is the most poisonous spider in the world, there is no proof for this statement. It is thought that this rumor may have been started because P. phalangioides kill and eat other spiders, including the Redback Spiders, whose venom can be fatal to humans. But to be able to kill the Redback spider, the P. phalangioides need only to have a quicker bite, not be more venomous (Crew, Haudon).

For More Information

Find Pholcus phalangioides information at

Contributors

Nancy Shefferly (editor), Animal Diversity Web Staff.

Anna Ferrick (author), Southwestern University, Stephanie Fabritius (editor), Southwestern University.

References

Emerton, J. 1902. The Common Spiders of the United Stateds. Boston, U.S.A., and London: Ginn & Company, Publishers.

Foelix, R. 1982. Biology of Spiders. Cambridge, Massachusets, and London, England: Harvard University Press.

Jackman, J. 1997. A Field Guide ot Spiders & Scorpions of Texas. Houston, TX: Gulf Publishing Company.

Stüber, K. 1999. "Biology of *Pholcus phalangioides*" (On-line). Accessed October 2, 2001 at http://caliban.mpiz-koeln.mpg.de/~stueber/essays/pholcus/pholcus_phalangioides.html.

Uhl, G. November 1998. Mating Behavior in the Cellar Spider, Pholcus phalangioides. Animal Behavior, Volume 56 Issue 5: 1155-1159.

To cite this page: Ferrick, A. 2002. "Pholcus phalangioides" (On-line), Animal Diversity Web. Accessed February 12, 2012 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Pholcus_phalangioides.html

Disclaimer: The Animal Diversity Web is an educational resource written largely by and for college students. ADW doesn't cover all species in the world, nor does it include all the latest scientific information about organisms we describe. Though we edit our accounts for accuracy, we cannot guarantee all information in those accounts. While ADW staff and contributors provide references to books and websites that we believe are reputable, we cannot necessarily endorse the contents of references beyond our control.

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