By Dr. Barbara Lundrigan and Marie Mueller
Geographic Range
Marmota camtscatica is found in Eastern Siberia (Steppan et al. 1999). Populations have been recorded in the mountains of northeastern Siberia from Lake Baikal to Kamchatka (Nowak 1991; Zimina and Gerasimov 1973).
Biogeographic Regions:
palearctic
(native
).
Habitat
Marmota camtschatica lives in extremely severe and variable conditions. Individuals live in mountainous areas (Macdonald 2001). Within the mountains, some live in forestless, alpine areas with permafrost and rocky areas, called goltzi, some live in the meadow-steppes on the warm south-facing slopes, some live in alpine meadows with Siberian dwarf pine (Pinus pumila), and still others inhabit ocean-facing volcanic slopes. While M. camtschatica is adapted to cold deserts and the harsh conditions at high elevations, it is not adapted to drought or dense forest (Zimina and Gerasimov 1973).
Terrestrial Biomes:
tundra
; taiga
; desert or dune
; mountains
.
Physical Description
(8.8 to 17.6 lbs; avg. 13.2 lbs)
(20.87 to 28.74 in; avg. 24.8 in)
Black-capped marmots are characterized by a large body size (Armitage 1999). They range from 53-73cm in length and 4-8kg in weight (Macdonald 2001).
They have black-tipped guard hairs, which provide excellent insulation in their cold environment (Steppan et al. 1999). They also have a bushy tail approximately half of their body length. These marmots have a plantigrade stance and walk on their entire foot. They have claws on their feet rather than nails.
Incisors are ever-growing and are self-sharpening. The cheekteeth are fan-shaped.
The eyes are placed on the side of the head and are surrounded by a light ring. Vibrissae are present on the head, feet, and legs (Macdonald 2001).
Some key physical features:
endothermic
; bilateral symmetry
.
Reproduction
Breeding occurs from March to May.
The single breeding pair is monogamous (Armitage 1999).
Mating systems:
monogamous
; cooperative breeder
.
Females start reproduction as soon as they emerge from hibernation so that the young have a sufficient amount of time to grow and prepare for hibernation the following fall(Armitage 1999). Females may even mate, start gestation, and give birth before they emerge from hibernation (Macdonald 2001). Fat resources are necessary for successful reproduction (Armitage 1999). Since females cannot accumulate enough resources to support annual reproduction, female M. camtschatica will skip reproduction and reproduce only every other year (Macdonald 2001). During lactation, females lose body mass (Armitage 1999). There is a single mating season in which 1-9 pups are born in April to June after 30-32 days of gestation (Nowak 1991).
Key reproductive features:
iteroparous
; seasonal breeding
; gonochoric/gonochoristic/dioecious (sexes separate); sexual
; fertilization
(internal
); viviparous
.
The young are born into an extended family group and are nursed by their mother in their natal burrow. Alloparenting by other members of the extended family occurs during hibernation (Armitage 1999).
Parental investment:
altricial
; male parental care
; female parental care
.
Lifespan/Longevity
Marmota camtschatica lives, on average, 4-5 years, with a maximum of 13-14 years. Females are known to live longer than males (Macdonald 2001).
Behavior
Marmota camtschatica individuals are diurnal and fossorial (Armitage 1999). They dig tunnels up to 113m in length (Nowak1991), though the average length is 10-70m (Zimina and Gerasimov 1973). They live in family groups made up of a breeding pair, subordinate adults, yearlings, and young (Armitage 1999). Black-capped marmots are the largest true hibernators. Fat stores are built up for hibernation. They hibernate in a common hibernaculum for social thermoregulation (Armitage 1999, Nowak 1991). Anywhere from 10-15 individuals may hibernate in one burrow (Zimina and Gerasimov 1973). Those individuals that disperse do so when they reach a body mass index (MI) of 0.5, which usually takes more than one year. While some females disperse, dispersal is primarily male-biased (Macdonald 2001; Armitage 1999).
Key behaviors:
motile
; sedentary
; social
; dominance hierarchies
.
Food Habits
Marmota camtschatica eat primarily green vegetation, especially plants that are in the early stages of growth (Nowak 1991; Zimina and Gerasimov 1973). They may also take fruits, grains, forbs, legumes and occasional insects (Nowak 1991). Marmota camtschatica do not store food (Armitage 1999).
Animal Foods:
insects.
Plant Foods:
leaves; seeds, grains, and nuts; fruit.
Predation
Black-capped marmots are only active during the summer months. They seek refuge from predators in their burrows and by being alert for predators and the alarm calls of conspecifics. Many medium to large carnivores may take black-capped marmots, including wolves and eagles.
Ecosystem Roles
Through their burrowing activities and herbivory black-capped marmots act to recycle and redistribute nutrients in the ecosystems in which they live. They also act as prey species for larger predators.
Economic Importance for Humans: Negative
Black-capped marmots inhabit regions that are sparsely populated by humans, there are no known adverse affects.
Economic Importance for Humans: Positive
Black-capped marmots are exploited as a source of fur (Steppan et al. 1999). They are important game animals, are important in epidemiological studies as vectors of diseases such as plague, and are important in paleontological research as they have left an extensive fossil record (Zimina and Gerasimov 1973).
Ways that people benefit from these animals:
food
; research and education.
Conservation Status
IUCN Red List: [link]:
Lower Risk - Least Concern.
US Federal List: [link]:
No special status.
CITES: [link]:
No special status.
Black-capped marmots are not currently recognized as threatened.
Other Comments
Marmota camtschatica provide biogeographic information about the faunal interchange across Beringia during the Plio-Pleistocene period (Steppan et al. 1999). The black-capped marmot is a good species in which to study the evolution of sociality (Armitage 1999).
For More Information
Find Marmota camtschatica information at
Contributors
Dr. Barbara Lundrigan
(author), Michigan State University. Marie Mueller (author), Michigan State University.

