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Lucanus elaphus


By Scott Teakell

Kingdom: Animalia
Class: Insecta
Order: Coleoptera
Family: Lucanidae
Genus: Lucanus
Species: Lucanus elaphus

Geographic Range

The giant stag beetle, also known as the elephant stag beetle, can be found in the woodlands of North America ranging from Virginia and North Carolina to the northeastern United States. (Arnett, Jr., Ph. D., 1985; Milne and Milne, 1980)

Biogeographic Regions
nearctic (Native )

Habitat

These beetles live and reproduce in damp, rotting wood. (Arnett, Jr., Ph. D., 1985; Drees, Ph. D. and Jackman, Ph. D., 1998; Milne and Milne, 1980)

Habitat Regions
temperate

Terrestrial Biomes
forest

Physical Description

Range length
30 to 60 mm
(1.18 to 2.36 in)

Adult males are 45-60 mm long; females are smaller, around 30-35 mm. This species has a slender, elongated form, with a somewhat flattened back. Adults are reddish-brown, shiny, and have black antennae and legs. Males have a crest above their eyes and a wide head. They are distinguished by their giant antlerlike jaws that may be as long as the head and thorax combined. These jaws have small forked teeth along the inner edge. The females have a narrower head than thorax and much smaller jaws than males. Female elytra are lightly punctate. Giant stag beetles have the segments of their antennae separated rather than compacted like scarab beetles do.

Larvae are white and grub-like, and are characterized by the absence of a 6th segment in the leg. (Arnett, Jr., Ph. D., 1985; Arnold and Drew, 1987; Drees, Ph. D. and Jackman, Ph. D., 1998; Klots and Klots, No Publishing Year; Milne and Milne, 1980; O'Toole, 1986)

Other Physical Features
ectothermic ; heterothermic ; bilateral symmetry

Sexual Dimorphism
male larger; ornamentation

Development

Giant stag beetle larvae hatch from eggs laid by females on suitable dead trees. They then eat and grow for several years in dead tree stumps. When fully-grown, the larvae pupate for seven to nine months, emerging the following June. After their emergence they live for about three to five weeks more. (Grzimek, 1972; Milne and Milne, 1980; Tweedie, 1973)

Development - Life Cycle
metamorphosis ; diapause

Reproduction

Males use their giant jaws to fight for access to females. Individual males try to control a dead tree or stump suitable for egg-laying, preventing other males from mating with the females arriving on the tree. Consequently one male usually mates with multiple females. (Milne and Milne, 1980)

Mating System
polygynous

Breeding season
In temperate climates, stag beetles breed in the summer months.

Female stag beetles lay their eggs on dead trees or stumps that will provide suitable food and protection for their offspring. In temperate climates, adults only live for a single breeding season.

Key Reproductive Features
semelparous ; seasonal breeding ; gonochoric/gonochoristic/dioecious (sexes separate); sexual ; fertilization (Internal ); oviparous

Eggs laid by female stag beetles are supplied with a small amount of nourishing yolk, but the beetle larvae hatch quickly, and receive no additional care. Male stag beetles do not care for their offspring.

Parental Investment
pre-fertilization (Provisioning, Protecting: Female)

Behavior

Males use thier impressive manibles in defense, as well as to compete with other males. Males compete with each other for access to females. Lucanus elaphus is attracted to lights at night. They can also sometimes be seen flying around dusk.

When males are challenged or forced to defend themselves they rear up using their forelegs and spread their jaws. This stance is mainly a bluff though, as their jaws can only pinch rather than inflict a painful bite.

Both males and females have difficulty getting upright if overturned because of their top-heavy heads and flattened backs.

Adults and larvae can be found in large colonies in burrows and rotted out logs. Adults can make noise by rubbing wing-covers or their legs together. (Burton, et al., 1968; Cottam, Ph. D. and Zim, Ph. D., 1956; Milne and Milne, 1980)

Key Behaviors
flies; crepuscular ; motile ; territorial ; colonial

Communication and Perception

Communication Channels
visual ; tactile ; chemical

Perception Channels
visual ; tactile ; vibrations ; chemical

Food Habits

Adult elephant stag beetles, like most stag beetles, feed on sugary liquid foods, mainly sap leaking from wounded trees, aphid "honeydew" secretions, and ripe fruit. They cannot chew food. The larvae feed on wet, decaying wood, probably getting nutrition from the wood and the fungi and microbes that are decomposing it. (Milne and Milne, 1980; O'Toole, 1986)

Plant Foods
wood, bark, or stems; fruit; sap or other plant fluids

Other Foods
fungus; microbes

Economic Importance for Humans: Positive

These large beetles are collected and raised by hobbyists. In the wild they can be important agents of wood decomposition.

Economic Importance for Humans: Negative

Though startling if found unexpectedly, these big beetles have no significant adverse effects on humans. They can pinch hard if handled carelessly, but only bite in self-defense. (Drees, Ph. D. and Jackman, Ph. D., 1998)

Negative Impacts
injures humans (bites or stings)

Conservation Status

IUCN Red List [Link]
No special status

US Federal List [Link]
No special status

CITES [Link]
No special status

This species is uncommon, but not believed to be in need of special conservation efforts.

For More Information

Find Lucanus elaphus information at

Contributors

Sara Diamond (editor), Animal Diversity Web Staff.

Scott Teakell (author), Southwestern University, Stephanie Fabritius (editor), Southwestern University.

References

Arnett, Jr., Ph. D., R. 1985. American Insects: A Handbook of the Insects of America North of Mexico. New York: Van Nostrand Reinhold Company.

Arnold, D., W. Drew. 1987. "The Stag Beetles of Oklahoma(Coleoptera: Lucanidae)" (On-line). Accessed February 18, 2001 at http://digital.library.okstate.edu/OAS/oas_htm_files/v67/p27_29nf.html.

Borror, D., R. White. 1970. A Field Guide to Insects: America North of Mexico. Boston: Houghton Mifflin Company.

Burton, J., I. Yarrow, A. Allen, L. Parmenter, I. Lansbury. 1968. The Oxford Book of Insects. Oxford: Oxford University Press.

Cottam, Ph. D., C., H. Zim, Ph. D.. 1956. Insects: A Guide to Familiar American Insects. New York: Western Publishing Company, Inc..

Drees, Ph. D., B., J. Jackman, Ph. D.. 1998. A Field Guide to Common Texas Insects. Houston, TX: Gulf Publishing Company.

Grzimek, D. 1972. Grzimek's Animal Life Encyclopedia Vol.2 Insects. England: Van Nostrand Reinhold Company.

Klots, A., E. Klots. No Publishing Year. Living Insects of the World. Garden City, New York: Doubleday and Company Inc..

Milne, L., M. Milne. 1980. The Audubon Society Field Guide to North American Insects and Spiders. New York: Alfred A. Knopf, Inc..

O'Toole, C. 1986. The Encyclopedia of Insects. New York: Equinox (Oxford) Ltd..

Tweedie, M. 1973. All Color Book of Insects. London: Octopus Books Limited.

To cite this page: Teakell, S. 2001. "Lucanus elaphus" (On-line), Animal Diversity Web. Accessed February 13, 2012 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Lucanus_elaphus.html

Disclaimer: The Animal Diversity Web is an educational resource written largely by and for college students. ADW doesn't cover all species in the world, nor does it include all the latest scientific information about organisms we describe. Though we edit our accounts for accuracy, we cannot guarantee all information in those accounts. While ADW staff and contributors provide references to books and websites that we believe are reputable, we cannot necessarily endorse the contents of references beyond our control.

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