By Alan Vu
Geographic Range
The natural distribution of European hares includes Great Britain and western Europe, east to through the Middle East to Central Asia (Lincoln, 1974; Broekhuizen and Maaskamp, 1980; Caillol and Meunier, 1989; Poli et al., 1991). They have been introduced by humans to several other continents. In Canada, Lepus europaeus is found in southern Ontario, around the Great Lakes, and south of the Canadian Shield. It has failed to spread further north. In the United States, European hares are now found in the north-eastern states and around the Great Lakes (Hall and Kelson, 1959). They have also been introduced to areas of South and Central America (Bonino and Montenegro, 1997) and Australia. (Bonino and Montenegro, 1997; Broekhuizen and Maaskamp, 1980; Caillol et al., 1988; Hall and Kelson, 1959; Lincoln, 1974; Poli et al., 1991)
Biogeographic Regions:
nearctic
(introduced
); palearctic
(native
); neotropical
(introduced
); australian
(introduced
).
Habitat
European hares prefer open fields and pastures bordered by hedgerows and woodlots, often around agriculture fields and crops. They live in shallow forms; clumps of grass, weeds, or bush (Peterson, 1966; Bansfield, 1974; William and Whitaker, 1943). (Bansfield, 1974; Hamilton and Whitaker, 1943; Peterson, 1966)
These animals are found in the following types of habitat:
temperate
; terrestrial
.
Terrestrial Biomes:
savanna or grassland
.
Other:
agricultural
.
Physical Description
(6.6 to 11 lbs)
(23.62 to 29.53 in; avg. 26.77 in)
Total length: 600-750 (average 680) mm; Ear length, from notch: 94-102 (av. 98) mm; Tail: 72-110 (av. 95) mm; Hind foot: 142-161 (av. 151) mm; Skull length: 96-104 (av. 100) mm; Skull width: 44-51 (av. 47.3) mm (Peterson, 1966; Hall and Kelson, 1959). They have long ears with black tips and which are greyish white inside. The pelage is yellowish-brown to greyish-brown, with a greyish-white underbody. The face is brown, with eye rings. The tail is black on the top and white on the bottom. In winter, L. europaeus doesn't change its pelage to white, but does become slightly more grey (Peterson, 1966; Bansfields, 1974; Dragg, 1974). There is no noted sexual dimorphism. The skull features short, broad, heavy nasal bones, and prominent anterior and posterior lobes of the supraorbital processes. It also often has a prominent subcutaneous process of the lacrimal bone, projecting from the anterior wall of the orbit (Bansfield, 1974). (Bansfield, 1974; Dragg, 1974; Hall and Kelson, 1959; Peterson, 1966)
Some key physical features:
endothermic
; homoiothermic; bilateral symmetry
.
Sexual dimorphism:
sexes alike.
Reproduction
There are two to several litters a season.
The breeding season for L. europaeus is between midwinter (January/February) and midsummer.
The breeding season for L. europaeus is between midwinter (January/February) and midsummer. The gestation period is between 30 and 42 days (Bansfield, 1974; Peterson, 1966). There is a high in-utero reabsorbtion rate; 7% in the spring to 25% in the autumn (Bansfield, 1974). Litter size varies between 1 and 8, the average being 3 to 5 (William and Whitaker, 1943; Bansfield, 1974). There are two to several litters a season. The weaning period is said to be about one month (Broekhuizen and Maaskamp, 1980; Bansfield, 1974). The young, called leverets, reach sexual maturity at eight months to a year in age. During autumn, the male's gonads and reproductive tract are regressed and plasma levels of testosterone and luteinizing hormone are low. In females, luteinizing hormone basal levels are at a maximum in July, the end of the reproductive season (Caillol and Meunier, 1989). (Bansfield, 1974; Broekhuizen and Maaskamp, 1980; Caillol et al., 1988; Hamilton and Whitaker, 1943)
Key reproductive features:
iteroparous
; seasonal breeding
; gonochoric/gonochoristic/dioecious (sexes separate); sexual
; viviparous
.
Leverets are precocial at birth, with long and silky fur (Peterson, 1966). To protect leverets, the mother disperses them over a moderately wide area to avoid predation on the whole litter. The mother then makes the rounds to nurse them (Bansfield, 1974). (Bansfield, 1974; Peterson, 1966)
Parental investment:
precocial
; pre-fertilization (provisioning, protecting: female); pre-hatching/birth (provisioning: female, protecting: female); pre-weaning/fledging (provisioning: female).
Behavior
European hares are mainly solitary animals except during mating season. They are crepuscular and nocturnal, mostly foraging at night (between 7 p.m. and 7 a.m.). European hares remain active all year round. During the day they crouch in a depression called a 'form', partially concealed with their back showing (Bansfield, 1974). European hares posess an excellent sense of sight, smell, and hearing. Upon detection of a predator, European hares will run to escape, and can dodge and change direction quickly if needed. They are very fast and have been clocked at up to 35 mph (about 60 kph) running in a straight line. They will also dive into streams if needed as they are decent swimmers (William and Whitaker; Bansfield, 1974). (Bansfield, 1974; Hamilton and Whitaker, 1943)
Home Range
There is little evidence to suggest that L. europaeus stays within a restricted home range. (Bansfield, 1974; Hamilton and Whitaker, 1943)
Key behaviors:
terricolous; saltatorial
; nocturnal
; motile
; sedentary
; solitary
.
Communication and Perception
European hares are usually quiet animals. They make low grunts from time to time and "guttural" calls from the doe (female) to her leverets. It has been suggested that European hares grind their teeth as an alarm call. They also emit a shrill call when hurt or caught (Peterson, 1966; Bansfield, 1974). (Bansfield, 1974; Peterson, 1966)
Food Habits
European hares are herbivorous, eating grasses, herbs, and field crops during summer. During winter European hares feed on twigs, buds, shrub bark, small trees, and young fruit tree bark. They also commonly re-ingest their green, soft fecal pellets. This is known as coprophagia. Two or three adult L. europaeus can eat as much vegetation as one sheep (Banfield, 1974; William and Whitaker, 1943; Peterson, 1966). (Bansfield, 1974; Hamilton and Whitaker, 1943; Peterson, 1966)
Primary Diet:
herbivore
(folivore
); coprophage
.
Plant Foods:
leaves; wood, bark, or stems.
Other Foods:
dung.
Predation
- red foxes (Vulpes vulpes)
- wolves (Canis lupus)
- coyotes (Canis latrans)
- larger hawks (Buteo)
- large owls (Strigiformes)
- wild cats (Felis silvestris)
Known predators include red foxes, wolves, coyotes (in their introduced range in North America), wild cats, larger hawks, and owls (Bansfield, 1974). (Bansfield, 1974)
Anti-predator adaptations::
cryptic
.
Economic Importance for Humans: Negative
In some areas, such as Argentina, Australia and, to a lesser extent, North America, L. europaeus is a pest. The problem lies in its quick reproduction and devastation to agriculture, especially young apple orchards (Bonino and Montenegro, 1997; Bansfield, 1974; William and Whitaker, 1943). (Bansfield, 1974; Bonino and Montenegro, 1997; Hamilton and Whitaker, 1943)
Ways that these animals might be a problem for humans:
crop pest; causes or carries domestic animal disease
.
Economic Importance for Humans: Positive
European hares have become an important and challenging game animal, especially in North America. The meat is said to be white and delicious (William and Whitaker, 1943; Bansfield, 1974). (Bansfield, 1974; Hamilton and Whitaker, 1943)
Ways that people benefit from these animals:
food
.
Conservation Status
IUCN Red List: [link]:
Lower Risk - Least Concern.
US Federal List: [link]:
No special status.
CITES: [link]:
No special status.
European hares are widespread throughout Europe, where they are called common hares. European hares have done well in North America, with population numbers quickly rising to the current density. In Ontario population density has been as high as 100 per square mile, and has leveled to about 25 per square mile (Bansfield, 1974; Dragg, 1974). In recent decades there have been outbreaks of increased mortality due to disease, particularly in Europe. This syndrome includes acute hepatosis, enteritis, nephrosis, general jaundice, congestion, and hemorrhage of internal organs, and has been called European Brown Hare Syndrome (Poli et al., 1991). (Bansfield, 1974; Dragg, 1974; Poli et al., 1991)
Other Comments
Lepus europaeus is native to Europe and South Asia, and was introduced everywhere else it is presently found. For example, it was introduced to Ontario in 1912, from Germany, and in New York State in 1893, and has been a successful game animal ever since (Dragg, 1974; William and Whitaker, 1943). In other countries such as Australia and Argentina, L. europaeus is a huge pest, with introduction resulting in agricultural disaster (Dragg, 1974; Bonino and Montenegro, 1997). Other common names for the European hare: common hare, brown hare (Caillol and Meunier, 1989; Poli et al., 1991) (Caillol et al., 1988; Dragg, 1974; Hamilton and Whitaker, 1943; Poli et al., 1991)
For More Information
Find Lepus europaeus information at
Contributors
Alan Vu (author), University of Toronto.

