By Jill Grogan
Geographic Range
Onagers, Equus hemionus onager, are found from Mongolia to Saudi Arabia and as far north as southern Russia and Kazakhstan. Some also inhabit northwestern India and Tibet. They have been reintroduced in Mongolia and Iran. ("Iran Nature and Wildlife Magazine", 2004; "IUCN SSC Equid Specialist Group", 2003; Wilson and Reeder, 1993)
Biogeographic Regions:
palearctic
(introduced
, native
).
Habitat
The flat regions of the deserts and surrounding foothills are home to E. hemionus onager. These barren deserts are a harsh environment and receive very little rainfall each year. ("Iran Nature and Wildlife Magazine", 2004; "IUCN SSC Equid Specialist Group", 2003; Moehlman and Feh, 2004; Wilson and Reeder, 1993)
These animals are found in the following types of habitat:
temperate
; terrestrial
.
Terrestrial Biomes:
desert or dune
.
Physical Description
In comparison with other Asiatic wild asses, E. hemionus onager is slightly smaller with a paler coat. Onagers have a pale sandy-red colored coat with a light brown dorsal stripe. The dorsal stripe has two surrounding white strips that blend into the lighter colored hind quarters. In addition to the dorsal stripe, onagers also have a shoulder stripe. The flanks, back and underside of onagers are white. In the winter, the coat grows longer and turns grayer and the white parts become more defined. Males and females differ only slightly outwardly, with males being only slightly larger. Males stand 1.5 meters at the shoulder and are about 2 meters in length, weighing about 250 kilograms. ("Iran Nature and Wildlife Magazine", 2004)
Some key physical features:
endothermic
; bilateral symmetry
.
Sexual dimorphism:
male larger.
Reproduction
Onager mares breed every other year.
Onagers usually mate in mid-June.
A dominant stallion mates with females in the herd. In order to assure breeding status, stallions defend the territories that females move through, with dominant stallions defending the best territories. ("Iran Nature and Wildlife Magazine", 2004; "IUCN SSC Equid Specialist Group", 2003)
Mating systems:
polygynous
.
During mating season in mid-June, stallions fight each other for mating rights. Females have a short estrus period of 3 to 5 days. After a year-long gestation, mares leave the herd to give birth in a safe place. A single foal is born that stays with its mother for two years. After giving birth, both mare and foal rejoin the herd, where the mother protects her foal from danger. ("Iran Nature and Wildlife Magazine", 2004; Nowak, 1999)
Pregnancy in onagers lasts 365 to 368 days. Nowak (1999) reports that lactation in female Equus hemionus (a species of wild ass in which some authorities include onagers) nurse their young for between 1 and 1.5 years. It is reasonable to assume that onagers fall within this range. Young onagers become independent around two years of age. Females are sexually mature around the age of 2, but males, at least in E. hemionus, mature about a year later. ("Iran Nature and Wildlife Magazine", 2004; Nowak, 1999)
Key reproductive features:
iteroparous
; seasonal breeding
; gonochoric/gonochoristic/dioecious (sexes separate); sexual
; fertilization
; viviparous
.
All members of the genus Equus are fairly precocial at birth, and are able to run shortly after birth. During the first two years of a foal's life, it stays with its mother. Mares protect their foals against all dangers. Nursing lasts between 1.5 and 2 years. Although specific details on the relationship between maternal dominance status and status of offsping, it is reasonable to assume that this species is like other members of the genus. If E. hemionus onager is like other horses, maternal dominance status affects status of the young within the herd. The role of males in care of offspring has not been reported. ("Iran Nature and Wildlife Magazine", 2004; Nowak, 1999)
Parental investment:
precocial
; pre-fertilization (provisioning, protecting: female); pre-hatching/birth (provisioning: female, protecting: female); pre-weaning/fledging (provisioning: female, protecting: female); pre-independence (protecting: female); maternal position in the dominance hierarchy affects status of young.
Lifespan/Longevity
The maximum lifespan of E. hemionus onager is reported to be approximately 40 years. However, whether this information is based on wild or captive animals is not known. ("Iran Nature and Wildlife Magazine", 2004)
Behavior
Equus hemionus onager usually lives in herds, with the exception of older stallions, who can be found living alone. Mares live with their foals in herds made up entirely of other females and young. ("Iran Nature and Wildlife Magazine", 2004; "IUCN SSC Equid Specialist Group", 2003)
Data specific to onagers is difficult to find, because of their debated taxonomic status. However, it is reasonable to assume that these animals are similar to E. hemionus, since they are sometimes placed in the same species. With that in mind, the following information on E. hemionus is provided.
Herd size may vary. In E. hemionus, average herds contain between 10 and 20 animals, with one male and many females. However, in some ecological conditions, pressure from predators causes small single male groups to come together. (Nowak, 1999)
Onagers are reported to be crepuscular. However, E. hemionus is reported to be primarily diurnal, feeding during the day and bedding down at night, with some populations feeding at night. (Nowak, 1999)
Equus hemionus is able to reach top speeds of about 70 km/hour, and can run at a sustained speed of 50 km/hour. It is likely that onagers are similar in their ability to attain such speeds. (Nowak, 1999)
Home Range
Because of the harsh conditions that the desert presents, E. hemionus onager must stay within 20 kilometers of a water source. ("Iran Nature and Wildlife Magazine", 2004; Nowak, 1999)
Key behaviors:
cursorial; terricolous; crepuscular
; motile
; nomadic
; migratory
; solitary
; social
; dominance hierarchies
.
Communication and Perception
Equus hemionus onager has strong senses with an extremely keen sense of smell. Like other members of the genus Equus, onagers have vocal, tactile and chemical communication. In addition, visual signals may be important. ("Iran Nature and Wildlife Magazine", 2004; Nowak, 1999)
Food Habits
Onagers are herbivores that feed on the scarce plant life in the desert. Foods of these animals include grasses, bushes, herbs and foliage. Onagers receive most of their water from their food, but must remain close to a site of open water. Grazing time for onager sis usually during the cooler part of the day such as morning and evening. ("Iran Nature and Wildlife Magazine", 2004; "IUCN SSC Equid Specialist Group", 2003; Moehlman and Feh, 2004; Wilson and Reeder, 1993)
Plant Foods:
leaves; roots and tubers; wood, bark, or stems; seeds, grains, and nuts.
Predation
Equus hemionus onager has no natural predators other than humans. This species is being driven to extinction due to hunting, competition for food and water with livestock, and loss of natural habitat. Onagers have a well developed sense of smell and can detect potential predators, such as humans, from a far distance. Onagers are also very fast, with the ability to run 60 to 70 kilometers per hour over short distances, and 40 to 50 kilometers per hour for several hours at a time. ("Iran Nature and Wildlife Magazine", 2004; "IUCN SSC Equid Specialist Group", 2003; "UNEP-WCMC Species Database: CITES-Listed Species", 2004; Bennett, 1980; Moehlman and Feh, 2004; Wilson and Reeder, 1993)
Ecosystem Roles
Grazing by E. hemionus onager impacts vegetation communities in which they live. ("Iran Nature and Wildlife Magazine", 2004)
Economic Importance for Humans: Negative
Although there is no apparent direct negative economic impact that this species has on humans, it is possible that these wild asses compete with livestock for water and scarce food resources. ("Iran Nature and Wildlife Magazine", 2004; "IUCN SSC Equid Specialist Group", 2003)
Economic Importance for Humans: Positive
Onagers have been hunted by humans in the past for their flesh and hides. However, in 1971, E. hemionus onager became a protected species in Iran and hunting it is prohibited year-round. ("Iran Nature and Wildlife Magazine", 2004; "IUCN SSC Equid Specialist Group", 2003; Moehlman and Feh, 2004)
Ways that people benefit from these animals:
food
; body parts are source of valuable material.
Conservation Status
IUCN Red List: [link]:
Critically Endangered.
US Federal List: [link]:
Endangered.
CITES: [link]:
Appendix II.
IUCN estimates 144 onagers remaining with the rate of decline at 28% over the last three generations. ("IUCN SSC Equid Specialist Group", 2003; "UNEP-WCMC Species Database: CITES-Listed Species", 2004; Moehlman and Feh, 2004)
Other Comments
Onagers are reported to have a bad temperment, which makes them unsuitable as work-a-day domestic animals. However, the ancient Roman Legions are thought to have used these animals to pull their war machines. Onagers were previously considered a species, Equus onager, but have since been included as a subspecies of kulans, Equus hemionus. (AZA Taxon Advisory Group, unknown)
For More Information
Find Equus hemionus onager information at
Contributors
Jill Grogan (author), University of Alaska Fairbanks. Link Olson (editor, instructor), University of Alaska Fairbanks.
Nancy Shefferly (editor), Animal Diversity Web Staff.

