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Dactyloscopidae
Sand stargazers


By Monica Weinheimer and R. Jamil Jonna

Kingdom: Animalia
Phylum: Chordata
Subphylum: Vertebrata
Class: Actinopterygii
Order: Perciformes
Family: Dactyloscopidae
Members of this Family

Diversity

Dactyloscopids derive their common name, sand stargazers, from their eyes, which protrude from the tops of their heads, sometimes on stalks. Sand stargazers usually remain burrowed into the sand with only the eyes, along with the snout and sometimes the top of the head, uncovered. Members of this family occupy shallow warm waters and are carnivorous. They are small, cryptic fishes and little is known about their behavior or reproductive habits. There are nine genera in the family and about 41 species. (Böhlke and Chaplin, 1993; Nelson, 1994; Springer, 1998; Wheeler, 1985)

Geographic Range

Dactyloscopids, or sand stargazers, are found in tropical and warm temperate waters of the western hemisphere. The Atlantic Ocean (from the United States to Brazil) contains 17 species, and the Pacific Ocean (from the Gulf of California to Chile) contains 24. (Allen and Robertson, 1994; Böhlke and Chaplin, 1993; Nelson, 1994; Springer, 1998; Wheeler, 1985)

Biogeographic Regions
nearctic (Native ); neotropical (Native ); atlantic ocean (Native ); pacific ocean (Native )

Habitat

Dactyloscopids live buried in the sand with only the eyes, along with the snout and sometimes the top of the head, uncovered. They usually occupy shallow warm water at depths between two and 15 m, but one species, Gillellus semicinctus, has been found between five and 137 m. Some groups inhabit bare, open beaches in or behind the surge zone, but others are found exclusively in patches of sand that are near rocks, coral structures, or marl bottoms. A few species can be found in estuaries, and at least one enters fresh water. (Allen and Robertson, 1994; Böhlke and Chaplin, 1993; Wheeler, 1985)

Habitat Regions
temperate ; tropical ; saltwater or marine

Aquatic Biomes
benthic ; coastal

Other Habitat Features
estuarine ; intertidal or littoral

Physical Description

Sand stargazers (family Dactyloscopidae) are small fishes adapted to burrowing in the sand. Their eyes are situated on the top of the head, sometimes on stalks, and their nostrils are tubular. They have upturned mouths, and fingerlike projections (fimbriae) line the mouth and gill covers, keeping sand out of the gills and mouth. These fish maintain water flow through the gills with a branchiostegal pump. Sand stargazers are covered with cycloid scales. The dorsal fin, which contains 7-23 spines and 12-36 soft rays, may be divided or continuous. The pelvic fins have three thickened rays with tips that are free from the fin membrane. The largest dactyloscopids reach 17 cm, but most are less than six cm. They are colored to blend in with their sandy environment: most are whitish or drab brown, and many have red-tinged or dark mottling along the back and head. (Click here to see a fish diagram) (Allen and Robertson, 1994; Böhlke and Chaplin, 1993; Nelson, 1994; Springer, 1998; Wheeler, 1985)

Other Physical Features
ectothermic ; bilateral symmetry

Sexual Dimorphism
sexes alike

Development

No information was found on development in Dactyloscopidae.

Reproduction

No information was found on mating systems in Dactyloscopidae.

While no information was found on reproduction in Dactyloscopidae, blennies (which are in the same suborder Blennioidei) in general tend to lay relatively large eggs that adhere to a surface. In the case of sand stargazers, eggs may adhere to each other, as males carry them in two clumps under the pectoral fins. (Thresher, 1984)

Key Reproductive Features
sexual ; fertilization (External ); oviparous

Sand stargazers display a unique trait among fishes in that males guard eggs by carrying them in two balls, one in the “armpit” (axilla) of each pectoral fin. (Allen and Robertson, 1994; Böhlke and Chaplin, 1993; Helfman, et al., 1997; Wheeler, 1985)

Parental Investment
male parental care

Lifespan/Longevity

No information was found on lifespan of dactyloscopids.

Behavior

Sand stargazers burrow in the sand and remain there most of the time lying in wait for their prey. They delve into the sand using sinuous body and anal fin motions, and swimming movements of the pectoral fins. They may bury themselves completely or leave the eyes, snout or top of the head uncovered. One behavior engaged in by males is egg-guarding, accomplished by carrying a ball of eggs under each pectoral fin. (Allen and Robertson, 1994; Böhlke and Chaplin, 1993; Springer, 1998; Wheeler, 1985)

Communication and Perception

No information was found on communication in Dactyloscopidae. Because their eyes are placed on top of their heads, often on stalks, so as to remain uncovered by sand, it is logical to infer that they perceive their surroundings visually.

Perception Channels
visual

Food Habits

Sand stargazers are carnivorous, lying in wait under the sand to attack small fishes and invertebrates. (Allen and Robertson, 1994; Wheeler, 1985)

Primary Diet
carnivore (Piscivore , Eats non-insect arthropods)

Predation

Sand stargazers conceal themselves from predators by hiding under the sand. They are cryptically colored, so any exposed parts blend in well with the sand. (Böhlke and Chaplin, 1993; Springer, 1998; Wheeler, 1985)

Anti-predator Adaptations
cryptic

Ecosystem Roles

Sand stargazers occupy a specific habitat, that is, sandy bottoms of shallow warm waters, and in that setting impact the tiny fishes and invertebrates that serve as their prey. (Allen and Robertson, 1994; Wheeler, 1985)

Economic Importance for Humans: Positive

No information was found on human importance of Dactyloscopidae.

Economic Importance for Humans: Negative

No specific information was found concerning any negative impacts to humans.

Conservation Status

IUCN Red List of Threatened Species [Link]
Not Evaluated.

There is no known conservation threat to any members of Dactyloscopidae. (The World Conservation Union, 2002)

For More Information

Find Dactyloscopidae information at

Contributors

Monica Weinheimer (author), Animal Diversity Web.

R. Jamil Jonna (author), Animal Diversity Web.

References

Allen, G., D. Robertson. 1994. Fishes of the Tropical Eastern Pacific. Honolulu, HI: University of Hawaii Press.

Böhlke, J., C. Chaplin. 1993. Fishes of the Bahamas and Adjacent Tropical Waters. Wynnewood, PA: Published for the Academy of Natural Sciences of Philadelphia by Livingston.

Helfman, G., B. Collete, D. Facey. 1997. The Diversity of Fishes. Malden, MA: Blackwell.

Nelson, J. 1994. Fishes of the World – third edition. New York, NY: John Wiley and Sons.

Springer, V. 1998. Blennies. Pp. 216 in W Eschmeyer, J Paxton, eds. Encyclopedia of Fishes – second edition. San Diego, CA: Academic Press.

The World Conservation Union, 2002. "IUCN 2002" (On-line). IUCN Red List of Threatened Species. Accessed August 18, 2003 at http://www.iucnredlist.org/.

Thresher, R. 1984. Reproduction in reef fishes. Neptune City, NJ: T.F.H. Publications.

Wheeler, A. 1985. The World Encyclopedia of Fishes. London: Macdonald.

To cite this page: Jonna, R. 2003. "Dactyloscopidae" (On-line), Animal Diversity Web. Accessed May 31, 2012 at http://animaldiversity.ummz.umich.edu/site/accounts/information/Dactyloscopidae.html

Disclaimer: The Animal Diversity Web is an educational resource written largely by and for college students. ADW doesn't cover all species in the world, nor does it include all the latest scientific information about organisms we describe. Though we edit our accounts for accuracy, we cannot guarantee all information in those accounts. While ADW staff and contributors provide references to books and websites that we believe are reputable, we cannot necessarily endorse the contents of references beyond our control.

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