By Rodgers Eckhart
Geographic Range
Capra walie, or the Walia Ibex, is found exclusively in the mountains of northern Ethiopia. Nearly all of the remaining population resides along 25 kilometers of the northern escarpment in the Simien Mountains National Park (Massicot, 2001).
Habitat
(8200 to 14760 ft)
The Simien Mountains are characterized by huge gorges and gulleys, both of which carve out steep and jagged cliffs. Walia Ibex individuals inhabit only the high cliffs that rise above the lower elevated plateau, providing a potential risk of falling for careless individuals (Beyene, 2001)
At a high elevation and low-latitude, the cliff habitat is conducive to extremes in the seasonality of precipitation and daily temperature fluctuation.
The wet season runs from May until October and is correlated with abundant plant growth and diversity. During the dry season, foods in the form of grasses and shrubs dramatically disappear from most of the landscape (Dunbar, 1978).
At an average elevation of nearly 3500 meters, the Walia Ibex encounters tremendous temperature fluctuations from night to day. On a normal day, the temperature ranges from near freezing to more than 25 degrees Celsius. Despite the fluctuations in daily temperatures, seasonal differences in temperature are minimal due to Ethiopia's proximity to the equator (Nievergelt, 1990).
These animals are found in the following types of habitat:
tropical
; terrestrial
.
Terrestrial Biomes:
mountains
.
Physical Description
(176 to 275 lbs; avg. 225.5 lbs)
Like other members of the genus Capra, Walia Ibex are sexually dimorphic in many aspects of appearance. In overall size, adult females weigh about 80 kilograms, roughly 50-60% of their male counterparts, which can weigh up to 125 kilograms (McDonald, 1984). Horns are semi-circular in shape and are found in both sexes, but male horns are more massive, reaching 110 centimeters in length (Beyene, 2001). Key features of horns are random knots and age rings, both of which distinguish individuals in a population (Dunbar and Dunbar, 1981).
Both sexes of Walia Ibex have black and white markings on their legs and a gray-white underside. The dorsal area is colored chestnut-brown and is darker in males (Nievergelt, 1990). In the wild, females are lighter in color and very inconspicuous (Dunbar, 1978). At older ages, males develop both a black chest and a "beard," further distinguishing the sexes (Dunbar and Dunbar, 1981).
Perhaps as an adaptation to its mountainous environment, Walia Ibex hooves have sharp edges and concave undersides that improves their grip by allowing them to work as a "suction cup" (Beyene, 2001).
Some key physical features:
endothermic
; bilateral symmetry
.
Reproduction
February-June
The typical mating system in C. Walie is polygyny, with dominant males siring a disproportionate amount of offspring during the breeding season. These males, because of their large size and fighting experience, are able to monopolize females by obtaining exclusive access to overlapping female home ranges (Dunbar and Dunbar, 1981).
Mating systems:
polygynous
.
C. walie is distinguished from other ibex species in its ability to breed at all times of the year. This may be possible because of the lack of temperature seasonality in the tropical Simien Mountains, producing no environmental costs to individuals that breed year-round (Nievergelt, 1990). Nevertheless, most often the Walia Ibex mates during the rut season, from February until April (Nievergelt, 1990). Peak sexual activity between males and females is observed between the months of March and June, overlapping with the short rut season (Dunbar and Dunbar, 1981). C. walie individuals reach sexual maturity at the age of one (Massicot, 2001). Both sexes continue to grow, however, and age is correlated with body size (Dunbar and Dunbar, 1981).
Key reproductive features:
seasonal breeding
; gonochoric/gonochoristic/dioecious (sexes separate); sexual
.
Behavior
Walia Ibex are strictly crepuscular, only actively moving or feeding in early mornings and late evenings. During the middle of the day, individuals become very lethargic and tend to hide under the shade of dense brush away from the intense noon sun and any nearby predators (Dunbar, 1978).
Atypical of most mountain sheep, Walia Ibex females are more solitary than males outside of the breeding season. Instead, males form small groups with other males of similar age or size (Dunbar and Dunbar, 1981). This pattern of association is reversed during the rut season, with females forming nursery groups and males isolating themselves from one another in competition (Dunbar, 1978).
Fighting during the rut season occurs most frequently between males of similar age and size. This resulst from the fact that older males almost always win their fights with younger competitors. In fact, if the age difference between two males is significant, fighting will usually not take place at all (Dunbar and Dunbar, 1981).
Key behaviors:
crepuscular
; motile
; social
; dominance hierarchies
.
Food Habits
A "generalized herbivore," Walia Ibex is both a grazer and a browser (Massicot, 2001). In fact, it utilizes a wide variety of grass and shrub material in its diet. Although grazing accounts for a significant part of its diet, this species spends most of its time feeding browsing in the cover of dense shrubs (Dunbar, 1978).
Foods eaten include: grasses, herbs, shrubs, bushes, creepers and lichens.
Plant Foods:
leaves; lichens.
Predation
- hyenas (Hyaeninae)
Hyenas appear to be the only predators capable of killing an adult Walia Ibex. However, juveniles are at risk from a large variety of predators ranging from wildcats to foxes (Dunbar, 1978).
Conservation Status
IUCN Red List: [link]:
Critically Endangered.
US Federal List: [link]:
Endangered.
CITES: [link]:
No special status.
Over the last half-century, Walia Ibex has been considered one of the most endangered ungulates in the world (Nievergelt, 1990). Today, this species is deemed "critically endangered" primarily because of the very small population left in the wild. In fact, recent estimates predict the number of remaining individuals to be no greater than 400 (Massicot, 2001).
Pressures on the remaining population take the form of limited habitat and poaching. Although not as large a problem as was in the past, poaching still does occur to some degree inside the national park (Beyene, 2001). On another note, increasing the population numbers in the future will inevitably be difficult as estimates predict that Walia Ibex's current habitat can only handle around 2000 individuals (Beyene, 2001).
Other Comments
Today, C. Walie is the southernmost species of Ibex found in the world (Massicot, 2001).
For More Information
Find Capra walie information at
Contributors
Rodgers Eckhart (author), University of Michigan.
Ondrej Podlaha (editor), University of Michigan.

